MORE EXOTIC AND FEWER NATIVE PLANT SPECIES: RIVERINE VEGETATION PATTERNS ASSOCIATED WITH ALTERED SEASONAL FLOW PATTERNS

Natural flow regimes are important for sustaining riverine vegetation. The regulation of river flows to provide water for agriculture often results in changes to flow timing. This study assesses the impact of altered seasonal flow patterns on riverine flora. Within temperate Australia, we surveyed the vegetation of five lowland rivers, three of which have large dams that alter their seasonal flow patterns; the other two are unregulated. From four to six sites were selected on each river, and these were classified into three levels of regulation based on the extent to which the timing of their seasonal flow patterns were altered. Sites were surveyed in winter and the following summer. Permanent quadrats were also established at a number of the surveyed sites and resurveyed every 3 months. Of the 267 plant taxa identified, 145 were exotic (non‐native). More exotic taxa and fewer native taxa were associated with increasing level of seasonal flow inversion (regulation). In particular, greater numbers of short‐lived exotic terrestrial taxa and fewer native woody taxa were associated with increasing level of regulation. Some exotic woody species (e.g. willows) were more common in the unregulated rivers and may have life‐history traits favoured by the natural seasonal flow patterns of study area. Multivariate analyses showed that level of regulation had a significant effect on the overall composition of the riverine vegetation. Our results provide support for the hypotheses that flow regulation adversely affects native species diversity and increases the vulnerability of riparian zones to invasion by exotic species; however, these effects are dependent on plant species' life‐history strategies. Our study highlights the importance of natural seasonal flow patterns for sustaining native riverine plant communities. Flow management aimed at maintaining or restoring ecological values should consider seasonal flow patterns. Winter/spring flow peaks may be particularly important for the recruitment of native riverine plants, especially trees and shrubs, and reducing the extent of exotic annuals and grasses. Copyright © 2012 John Wiley & Sons, Ltd.     

ECOSYSTEM ENGINEERS MODULATE EXOTIC INVASIONS IN RIPARIAN PLANT COMMUNITIES BY MODIFYING HYDROGEOMORPHIC CONNECTIVITY

Patterns of native and exotic plant species richness and cover were examined in relation with ecosystem engineer effects of pioneer vegetation within the Mediterranean gravel bed river Tech, South France. The floristic composition was characterized according to two distinct vegetation types corresponding to two habitats with contrasted conditions: (i) open and exposed alluvial bars dominated by herbaceous communities; and (ii) islands and river margins disconnected from annual hydrogeomorphic disturbances and covered by woody vegetation. A significant positive correlation between exotic and native plant species richness and cover was observed for both vegetation types. However, significant differences in native and exotic species richness and cover were found between these two vegetation types. Higher values of total species richness and Shannon diversity were attained within the herbaceous vegetation type than within the woody type. These differences are most likely related to changes in local exposure to hydrogeomorphic disturbances driven by woody engineer plant species and to vegetation succession. A lower exotic species cover within the woody vegetation type than within the herbaceous type suggested an increase of resistance to invasion by exotic species during the biogeomorphic succession. The engineer effects of woody vegetation through landform construction resulted in a decrease of alpha (α) diversity at the patch scale but, in parallel, caused an increase in gamma (γ) diversity at the scale of the studied river segment. Our study corroborates recent investigations that support the theory of biotic acceptance of exotic species by native species at the local scale (generally <10 m²) within heterogeneous and disturbed environments. Furthermore, we suggest that in riparian contexts such as the River Tech exotic species trapp sediment at the same time as native species and thus contribute to the increase in ecosystem resistance during the biogeomorphic succession. Copyright © 2012 John Wiley & Sons, Ltd.     

Changes in the hydrological regime and invasions by plant species along riparian systems of the Adour River,France

Changes in the hydrological regime affect the phenomenon of invasion by plant species along riparian systems. The dynamics of exotic (non-native) and native species were examined at five sites that differed in exposure to hydrological disturbance (floods) during three consecutive years (medium, wet and dry years). When considering the disturbance gradient, exotic plants were favoured by direct exposure to floods (main channel) and by high flood frequencies. The response to year to year changes in hydrology was rapid for both native and exotic communities. However, the exotic plants responded more rapidly and were favoured by a dry year. A general framework including human and natural factors involved in invasions by exotic plants along rivers is presented.     

Effects of stream flow intermittency on riparian vegetation of a semiarid region river (San Pedro River,Arizona)

The San Pedro River in the southwestern United States retains a natural flood regime and has several reaches with perennial stream flow and shallow ground water. However, much of the river flows intermittently. Urbanization‐linked declines in regional ground‐water levels have raised concerns over the future status of the riverine ecosystem in some parts of the river, while restoration‐linked decreases in agricultural ground‐water pumping are expected to increase stream flows in other parts. This study describes the response of the streamside herbaceous vegetation to changes in stream flow permanence. During the early summer dry season, streamside herbaceous cover and species richness declined continuously across spatial gradients of flow permanence, and composition shifted from hydric to mesic species at sites with more intermittent flow. Hydrologic threshold values were evident for one plant functional group: Schoenoplectus acutus, Juncus torreyi, and other hydric riparian plants declined sharply in cover with loss of perennial stream flow. In contrast, cover of mesic riparian perennials (including Cynodon dactylon, an introduced species) increased at sites with intermittent flow. Patterns of hydric and mesic riparian annuals varied by season: in the early summer dry season their cover declined continuously as flow became more intermittent, while in the late summer wet season their cover increased as the flow became more intermittent. Periodic drought at the intermittent sites may increase opportunities for establishment of these annuals during the monsoonal flood season. During the late summer flood season, stream flow was present at most sites, and fewer vegetation traits were correlated with flow permanence; cover and richness were correlated with other environmental factors including site elevation and substrate nitrate level and particle size. Although perennial‐flow and intermittent‐flow sites support different streamside plant communities, all of the plant functional groups are abundant at perennial‐flow sites when viewing the ecosystem at broader spatial and temporal scales: mesic riparian perennials are common in the floodplain zone adjacent to the river channel and late‐summer hydric and mesic annuals are periodically abundant after large floods. Copyright © 2005 John Wiley & Sons, Ltd.     

Soil seed banks of degraded riparian zones in southeastern Australia and their potential contribution to the restoration of understorey vegetation

Although soil seed banks are understood to be integral to the vegetation dynamics and restoration of many ecosystems, little is known of their role in riparian zones. In this study, we investigated soil seed banks of riparian zones of contrasting condition in an agricultural landscape and evaluated their potential to influence riparian restoration. We examined the composition and structure of germinable soil seed banks along lateral gradients from stream channels in both cleared and wooded riparian zones of three lowland creeks within the Goulburn Broken catchment in temperate southeastern Australia. Environmental correlates of soil seed bank characteristics and similarity to extant vegetation were also examined. We found an abundant and species‐rich soil seed bank mostly comprising propagules of perennial rushes and sedges and annual and perennial grasses with many species of annual forbs. While the majority of identifiable germinants and species were native, exotic species were common at all locations. Soil seed bank composition was relatively homogeneous among streams and along lateral gradients from the channel. Riparian condition (i.e. cleared or wooded), however, had a strong influence on soil seed bank composition and structure with cleared reaches containing more species, more germinable annual grasses and higher total numbers of germinable seeds. Soil seed bank composition was correlated with site openness suggesting that extant vegetation structure plays an important role in soil seed bank dynamics. Recruitment from the in situ soil seed bank will help restore only some components of the riparian plant community and may hinder restoration by introducing undesirable species. Copyright © 2008 John Wiley & Sons, Ltd.     

Reptile community responses to native and non‐native riparian forests and disturbance along two rivers in Arizona

Aridland riparian forests are undergoing compositional changes in vegetation and wildlife communities due to altered hydrology. As flows have been modified, woody vegetation has shifted from native‐tree dominated to non‐native and shrub encroached habitats. Squamate vertebrates such as lizards and snakes are important food web links in riparian ecosystems of the Sonoran Desert. However, little is known about how these communities might respond as riparian forests transition from native tree dominated habitats to open xeroriparian woodlands. We used pitfall arrays deployed across three types of riparian forest to document reptile community patterns, measure vegetation, and produce species‐habitat models. Riparian forests differed on the basis of habitat composition and physiognomy. Two types, cottonwood‐willow (Populus‐Salix) and mesquite (Prosopis) stands, were characterized by high woody species richness. The third type, non‐native saltcedar (Tamarix) stands, had high densities of woody debris and greater canopy coverage. Results show that lizards were common and abundances greatest in cottonwood‐willow, especially for arboreal species. Species‐habitat models for three of five lizard species indicated a negative association to saltcedar‐invaded habitat and no species appeared to select saltcedar‐dominated habitat. Mesquite was an intermediate habitat between upland and riparian, and supports high species diversity. A wildfire in the cottonwood‐willow forest disproportionately affected abundance of ground‐foraging whiptail (Aspidoscelis) lizards; whereas, abundance of arboreal spiny (Sceloporus) species was unchanged. Expected drivers from climate and water use could transition cottonwood forests to other woody‐dominated types. Our results suggest that mesquite woodlands would provide higher quality habitat for riparian reptiles compared to non‐native saltcedar stands.     

Abundance of Invasive,Non‐Native Riparian Herbs in Relation to River Regulation

River regulation is associated with vegetation encroachment and invasions of some non‐native species in the semi‐arid west. Shifts in the abundance of native and non‐native woody riparian species are an interplay of regulation, life history traits and an array of flow and physical environmental variables. We sought to compare plant densities and per cent cover of several invasive species over two time periods in a paired river study, contrasting three different degrees of regulation along reaches of the Green and Yampa rivers in Colorado and Utah, USA. We censused patches of non‐native plants and recorded per cent cover in quadrats along 171 river km. The upper Green (10.1 patches ha^?1) had the highest invasive plant patch density followed by the lower Green (4.4 per ha) and the Yampa (3.3 per ha). Invasive species were present in 23%, 19% and 4% of sample quadrats, and an average of 0.28, 0.22 and 0.04 invasive species detected per square metre was recorded along the upper Green, lower Green and Yampa Rivers, respectively. Most species had significantly (p ≤ 0.02) higher percent cover on the upper Green than either or both the lower Green and the Yampa River. Whereas the less regulated river reaches maintain lower densities of invasive species than the most regulated reach, long‐term persistence of this pattern is still in question as some species patches showed notable increases on the Yampa and lower Green Rivers from 2002–2005 to 2010–2011. Although invasion is enhanced by flow regulation, life history traits of some species suggest invasion is likely, regardless of flow regulation. Published 2015. This article is a U.S. Government work and is in the public domain in the USA     

Rodent use of anthropogenic and ‘natural’ desert riparian habitat,lower Colorado River,Arizona

The role of native trees, Fremont cottonwood (Populus fremontii) and Goodding willow (Salix gooddingii), in structuring the riparian small mammal assemblage on rivers in the American desert southwest is unclear. It is unknown, for example, whether these trees directly or indirectly provide the food or shelter necessary for the presence of any species. Because of the rapid and widespread decline of gallery forest, due in part to river regulation, the retention of remnant stands and replacement of lost stands are major regional conservation issues. To elucidate small mammal–forest linkages, we compared patterns of macrohabitat use among terrestrial small mammals at two rehabilitated and one unmanipulated alluvial floodplain site along the highly regulated lower Colorado River. We also compared current patterns to the Colorado River faunal associations Joseph Grinnell documented in 1910, prior to significant flow regulation. We used grid‐based, capture‐mark‐recapture techniques at two revegetation sites, each a mosaic of six distinct macrohabitats, including planted cottonwood/willow, to associate species with specific macrohabitats. We also trapped a ‘reference’ grid containing naturally regenerating cottonwood and willow at a site on the lower Bill Williams River floodplain. Despite very poor development of cottonwood plantings at one of the revegetated sites, each supported at least nine species and harbored all seven species that Grinnell associated with areas flooded nearly every year. The set of species Grinnell associated with cottonwood/willow stands (Peromyscus maniculatus, Reithrodontomys megalotis, and Sigmodon arizonae) was trapped at both revegetation sites but entirely absent at the reference site. The Bill Williams site may be inaccessible to Sigmodon, but the absence of the other two species is probably a consequence of differences in floodplain structure and functioning among the study sites as well as between the Bill Williams site and historic Colorado River riparian areas. Our data suggest the richness of the native lower Colorado River riparian small mammal assemblage is unrelated to the presence or absence of cottonwood/willow trees, but does depend in part upon the presence or absence of dense herbaceous vegetation. Resource managers attempting to rehabilitate degraded desert riverine ecosystems need to consider understory as well as overstory plant species in revegetation efforts. Copyright © 1999 John Wiley & Sons, Ltd.     

Effects of Urban Sprawl on Riparian Vegetation: Is Compact or Dispersed Urbanization Better for Biodiversity?

Compact urbanization is the main strategy for sustainable urban development. However, it is yet unclear whether compact urban forms are ecologically more favourable than dispersed ones. In this paper, we studied the effects of urban sprawl on the riparian vegetation condition in one of the most degraded watersheds in the Buenos Aires metropolitan area, Argentina. We conducted random sampling of the riparian vegetation at sites along streams in the basin and assessed urban indicators at the reach and sub‐watershed scales for each of those sites in a geographic information system: urban area, impervious surface, population density and two landscape metrics of dispersion. The indicators assessed explained a high proportion of the variability of the vegetation response variables, thereby confirming the importance of urban sprawl pressure in shaping riparian communities in fluvial ecosystems. Dispersed urbanization had more positive than negative effects on the vegetation in the study area. Riverbanks associated with dispersed urbanization had more plant species, including exotics, when urban sprawl was assessed at the local scale. At the sub‐watershed scale, dispersed urbanized areas were richer in native plants and most of the functional groups, and poorer in exotic species. The model of the compact city, including bio‐corridors along watercourses, has been proposed for the Buenos Aires conurbation process for the next decades. Our results showed that the quality of existing river corridors across the compact matrix was not desirable and best practices for redesigning a more sustainable landscape structure are necessary, including the restoration of habitats for wetland species. Copyright © 2017 John Wiley & Sons, Ltd.     

Ecological strategies successfully predict the effects of river floodplain rehabilitation on breeding birds

To improve the ecological functioning of riverine ecosystems, large‐scale floodplain rehabilitation has been carried out in the Rhine–Meuse Delta since the 1990s. This paper evaluates changes in abundance of 93 breeding bird species over a period of 10 years in response to rehabilitation, by comparing population changes in 75 rehabilitated sites with 124 non‐rehabilitated reference sites. Such quantitative, multi‐species, large‐scale and long‐term evaluations of floodplain rehabilitation on biodiversity are still scarce, particularly studies that focus on the terrestrial component. We try to understand the effects by relating population trends to ecological and life‐history traits and strategies of breeding birds. More specifically, we try to answer the question whether rehabilitation of vegetation succession or hydro‐geomorphological river processes is the key driver behind recent population changes in rehabilitated sites. Populations of 35 species have significantly performed better in rehabilitated sites compared to non‐rehabilitated floodplains, whereas only 8 have responded negatively to rehabilitation. Differences in effects between species are best explained by the trait selection of nest location. Reproductive investment and migratory behaviour were less strong predictors. Based on these three traits we defined eight life‐history strategies that successfully captured a substantial amount of variation in rehabilitation effects. We conclude that spontaneous vegetation succession and initial excavations are currently more important drivers of population changes than rehabilitation of hydrodynamics. The latter are strongly constrained by river regulation. If rehabilitation of hydro‐geomorphological processes remains incomplete in future, artificial cyclic floodplain rejuvenation will be necessary for sustainable conservation of characteristic river birds. Copyright © 2010 John Wiley & Sons, Ltd.     

DRIVERS OF RIPARIAN TREE INVASION ON A DESERT STREAM

Understanding mechanisms of exotic species' invasions is essential to managing riparian landscapes throughout the world. In the southwestern USA, the two most dominant invaders of riparian habitats are the exotic tree species tamarisk (Tamarix ramosissima, Tamarix chinensis, and their hybrids) and Russian olive (Elaeagnus angustifolia). These plants were introduced around 1900, and their success may be facilitated by river regulation, river channel changes, and precipitation patterns. We hypothesized that riparian invasion in Canyon de Chelly, Arizona, was initiated by a change point event such as plantings, dam construction, or channel incision and that establishment near a change point was tied to flood events. We aged tamarisk, Russian olive, and native cottonwood trees from study sites in Canyon de Chelly and used tree ring analysis to determine the year of establishment and the elevation of the germination point relative to the channel. We used Bayesian Poisson regression and information theoretics to identify change points and precipitation variables driving annual tree establishment. We found that most tamarisk and Russian olive trees established in the late 1980s, and most cottonwoods established in 1930‐1950 and 1980‐2000. Regression models indicated that change points occurred in 1983 for Russian olive and 1988 for tamarisk, and precipitation was important for establishment. Although plantings and river regulation probably played a role in tree invasion, our results suggest that these species required precipitation and stream channel change for widespread establishment in Canyon de Chelly. The factors driving riparian invasions may not be those often associated with degraded rivers, such as altered hydrographs and land management changes, thus requiring analyses of the full range of ecological and physical processes. Copyright © 2012 John Wiley & Sons, Ltd.     

DECLINE AND RECOVERY OF SMALL MAMMALS AFTER FLOODING: IMPLICATIONS FOR PEST MANAGEMENT AND FLOODPLAIN COMMUNITY DYNAMICS

Floodplains often are managed both for agriculture and as habitat for native species. On the Sacramento River, farmers have expressed concern that natural areas may harbour small mammals in greater abundance than occurs on farms, which could make them sources of pests to adjoining farmlands. Concern over this has generated some opposition to riparian restoration. To better understand the issue, we studied small mammals at agricultural sites, young and older riparian restoration sites and remnant natural areas through live trapping and visual surveys over three successive years. Significant flooding occurred during the second winter of the study. We found that young riparian restoration sites (3–4 years since planting) did not harbour agricultural pests (e.g. California ground squirrel, Botta's pocket gopher) in greater abundance than agricultural sites, with the exception of California voles. Although voles were common at young sites, they declined dramatically as the sites matured, and neither 12‐ to 15‐year‐old sites nor remnant riparian habitats had higher abundances of any small mammal pest species than agricultural sites. All species except the arboreal western gray squirrel experienced significant population declines following the flood and remained at depressed levels for at least a full year thereafter. Native species were not less susceptible to flooding than exotic species. California vole was the first species to recover. Our study suggests that frequent winter flooding may be desirable from an agricultural pest management perspective. It illustrates a valuable ecosystem service, small mammal pest control, which is provided by flooding, an important natural disturbance process. Copyright © 2011 John Wiley & Sons, Ltd.     

A ROADMAP FOR RIPARIAN INVASION RESEARCH

Riparian habitats provide numerous ecosystem services and support a diversity of ecological processes, yet they are environmentally sensitive landscapes susceptible to species invasion. As the global population grows, urban, exurban, suburban and rural development of riparian landscapes and dependence on the associated ecosystem services will increase. To preserve riparian systems, research should assess causes and consequences of riparian invasion and effective restoration measures across a diversity of biomes. This paper categorizes the types of riparian invasion research studies and investigates research trends in the riparian invasion literature. The analysis shows that research focuses on drivers and patterns of invasion is primarily focused on North America and Europe and is scant in high biodiversity biomes, such as the tropics and humid subtropics. Future population growth is predicted to occur in poorly researched regions. Future riparian invasion research should focus on regions of high biodiversity and identify consequences of invasion and restoration measures. Copyright © 2013 John Wiley & Sons, Ltd.     

Legacies of flood reduction on a Dryland river

The Bill Williams (Arizona) is a regulated dryland river that is being managed, in part, for biodiversity via flow management. To inform management, we contrasted riparian plant communities between the Bill Williams and an upstream free‐flowing tributary (Santa Maria). Goals of a first study (1996–1997) were to identify environmental controls on herbaceous species richness and compare richness among forest types. Analyses revealed that herbaceous species richness was negatively related to woody stem density, basal area and litter cover and positively related to light levels. Introduced Tamarix spp. was more frequent at the Bill Williams, but all three main forest types (Tamarix, Salix/Populus, Prosopis) had low understory richness, as well as high stem density and low light, on the Bill Williams as compared to the Santa Maria. The few edaphic differences between rivers (higher salinity at Bill Williams) had only weak connections with richness. A second study (2006–2007) focused on floristic richness at larger spatial scales. It revealed that during spring, and for the study cumulatively (spring and fall samplings combined), the riparian zone of the unregulated river had considerably more plant species. Annuals (vs. herbaceous perennials and woody species) showed the largest between‐river difference. Relative richness of exotic (vs. native) species did not differ. We conclude that: (1) The legacy of reduced scouring frequency and extent at the Bill Williams has reduced the open space available for colonization by annuals; and (2) Change in forest biomass structure, more so than change in forest composition, is the major driver of changes in plant species richness along this flow‐altered river. Our study informs dryland river management options by revealing trade‐offs that exist between forest biomass structure and plant species richness. Copyright © 2010 John Wiley & Sons, Ltd.     

Does species trait composition influence macroinvertebrate responses to irrigation water withdrawals: Evidence from the Intermountain West,USA

Irrigation water withdrawals are ubiquitous in arid and semi‐arid regions; however, predictable macroinvertebrate responses have not emerged. Inconsistent responses are frequently attributed to the amount of water withdrawn, while physicochemical and biological differences among systems remain unstudied. In this study, we sought to determine if species traits influence macroinvertebrate responses to water withdrawals. We sampled above and below 12 independent water diversions experiencing discharge reductions greater than 75% in central and eastern Oregon and Washington, USA. When considering all 12 rivers, we observed significant alterations to the hydrologic environment, in addition to increased algal standing stocks, silt cover and minimum temperatures; however, macroinvertebrate responses were limited to increased downstream densities, as indices and community composition did not significantly change. Functionally, species traits were more sensitive than structural indices; proportional abundance for 12 of 52 species traits significantly differed above and below all points of diversion. Trait changes suggest a shift in functional composition to compensate for hydrologic alterations and reduced habitat area. Grouping references sites (upstream) by species trait composition revealed a strong ambient gradient of increasing resistance and resilience from high to low elevations; however, we did not find evidence for differential susceptibility to water withdrawals among trait‐based reference groups. Though water withdrawals changed the representation of select species traits, frequencies of trait states most vulnerable to water withdrawals were too homogenously distributed among sites to elicit differential responses among trait‐based reference groups. At larger spatial scales or among disparate hydrologic regimes, grouping systems by species traits will likely reduce variability and offer a more general currency for detecting perturbation response relationships. Copyright © 2009 John Wiley & Sons, Ltd.     

Identifying the natural flow regime and the relationship with riparian vegetation for two contrasting western Australian rivers

The natural flow regime and the relationship between flows and riparian vegetation are described for sites on both the Blackwood River in south‐western Australia and the Ord River in north‐western Australia. Analysis of long‐term flow data showed the historic mean monthly river discharge for the Blackwood River is strongly seasonal and highly predictable with generally low variability each month. The Ord River showed a strong seasonality of flows with about 92% of the (total) yearly flow occurring between December and March. Flow variability was very high (e.g. coefficient of variation >100% for all months) but highly predictable, with this mostly attributed to low but constant dry‐season flows. Water depth, duration of flood events and the number of flood events per year show a significant correlation with aspects of the riparian vegetation within experimental vegetation plots. Results highlight the strong relationship between floristics, life form structure and population dynamics with stream hydrology. On the Blackwood River, species richness and cover of shrubs reduced with increased duration and frequency of flooding, while cover of exotic species and annual herbs increased with increased flooding. Germination of tree seedlings was not influenced by flood regime but size class of tree species increased with flooding frequency. On the Ord River, species richness was not influenced by flooding regime. However, cover of perennial grasses increased with flooding frequency whilst cover of shrubs decreased. There was no relationship between flooding and seedling establishment whilst tree size class decreased with increased flooding. The methods described here can be used to compare the response of different components of the riparian vegetation to different fluvial regimes (e.g. because of impoundment and abstraction). This technique can be expanded for the management of riparian zones and planning rehabilitation programmes. It may also be useful for improving the ecological knowledge base for setting environmental flows in regulated systems. Copyright © 2001 John Wiley & Sons, Ltd.     

Butterfly (Papilionoidea and Hesperioidea) assemblages associated with natural,exotic, and restored riparian habitats along the lower Colorado River,USA

Butterfly assemblages were used to compare revegetated and natural riparian areas along the lower Colorado River. Species richness and correspondence analyses of assemblages showed that revegetated sites had fewer biological elements than more natural sites along the Bill Williams River. Data suggest that revegetated sites do not provide resources needed by some members of the butterfly assemblage, especially those species historically associated with the cottonwood/willow ecosystem. Revegetated sites generally lacked nectar resources, larval host plants, and closed canopies. The riparian system along the regulated river segment that contains these small revegetated sites also appears to have diminished habitat heterogeneity and uncoupled riparian corridors. Revegetated sites were static environments without the successional stages caused by flooding disturbance found in more natural systems. We hypothesize that revegetation coupled with a more natural hydrology is important for restoration of butterfly assemblages along the lower Colorado River. Copyright © 1999 John Wiley & Sons, Ltd.