小型湖泊应用于地表水源热泵时的热承载能力分析

介绍了小型湖泊应用于地表水源热泵时的两种热承载能力计算方法——水面温度法和均匀混合法，并采用这两种方法计算了杭州小型湖泊应用于地表水源热泵时的热承载能力。计算结果表明：水面温度法较客观地反映了实际的排水混合及水温恢复过程，可信度更高。用水面温度法对7个代表性城市小型湖泊热承载能力进行分析的结果表明：夏季热承载能力在23.4～29.5 W/m2之间，冬季热承载能力在31.6～45.7W/m2之间；在设计这类系统时关键在于满足夏季热承载能力的约束。     

Water‐related bird assemblages in an urban pond ‘archipelago’: Winter patterns of bird species occurrence,abundance and richness

This study reports on the patterns of species occurrence, abundance and richness of a wintering water‐related bird assemblage in an ‘archipelago’ of 70 small artificial urban ponds (AUPs) embedded in a metropolitan landscape (Rome, central Italy). A total of 20 species in 26 AUPs were sampled. Only the largest AUPs (>0.1 ha) contained all these species, except for Gallinula chloropus. The highest total mean species abundance was observed in the largest ponds, with statistically significant differences evident among size classes. Two significant spatial thresholds in species abundance and richness were observed (between 0.01 and 0.1 ha; between 0.1 and 1 ha in size). The abundance of single species was correlated with their frequency of occurrence. Ponds in urban areas must be larger than 0.1 ha to host a rich winter assemblage of birds, with a further increase in richness noted with a surface area larger than 1 ha. The highest number of species was observed in the larger ponds (>1 ha). The species richness of each AUP is directly correlated to their size (log‐transformed species–area relationship: log S = 3.515 + 0.497 log A; R² = 0.76). Further research should be conducted to confirm these patterns and to implement information useful for planning and management of artificial ponds in urban areas for this purpose.     

The effects of water retention time and watershed features on the limnology of two tropical reservoirs in Brazil

Although reservoirs are similar to natural lakes in many respects, such driving forces as water retention time and watershed features can play important roles in the limnology of manmade lakes. With the goal of investigating how these factors influence the limnology of tropical reservoirs, physical and chemical variables were measured at four sampling sites in two reservoirs in southern Brazil, from June 2002 to June 2003. Funil Reservoir is located in one of the most-populated areas in the country, in the Paraíba do Sul river basin, which drains and drastically influences the water quality of the reservoir. In contrast, Lajes Reservoir is located in a well-preserved area, with its water retention time varying from six to 30 times longer than for Funil Reservoir. Funil Reservoir is a turbid (median euphotic zone = 4.3 m), eutrophic reservoir (median total phosphorus (TP) = 3.1 µm ), with a high phytoplankton biomass (median chlorophyll-a concentration = 10.0 µg L⁻¹). In contrast, Lajes Reservoir is a transparent (median euphotic zone = 9.2 m), mesotrophic water system (median TP = 1.0 µm ), with a low phytoplankton biomass (median chlorophyll-a = 1.9 µg L⁻¹). Both reservoirs were stratified during the summer months, but isothermy was only observed in Funil Reservoir. Because of its short water retention time, Funil Reservoir is a much more dynamic system than Lajes Reservoir, with a pronounced temporal pattern related to changes in its water column and its phytoplankton biomass. Spatial heterogeneity is more evident in Lajes Reservoir, mainly as a consequence of its location in a preserved area, long water retention time and the presence of net cages for fish culture in the waterbody. The typical spatial zonation found in reservoirs, related to nutrient sedimentation and light availability, however, is more evident in Funil Reservoir than in Lajes Reservoir. Despite the similarities between these two water systems, which are in the same geographical region with similar climate, and are comparable in size, the distinct watershed features and water retention time are responsible for marked differences between these reservoirs.     

Satellite altimetry for Indian reservoirs

Satellite radar altimetry has immense potential for monitoring fresh surface water resources and predicting the intra-seasonal, seasonal, and inter-annual variability of inundated surface water over large river basins. As part of the Preparation for the Surface Water and Ocean Topography mission scheduled for launch in mid-2022, the present study aimed to evaluate the performance of radar altimetry over the inland water bodies of India. The Joint Altimetry Satellite Oceanography Network (Jason) and Satellite with ARgos and ALtiKa (SARAL/AltiKa) data were used to derive the water levels of 18 major reservoirs in India by incorporating the geophysical and propagation corrections into the radar range. In situ gauge data were used to evaluate the performance of the altimetry-derived water level time series from 2008 to 2019. The results showed a strong correlation between Jason-2 and in situ data with the determination coefficient (R²) and root mean squared error (RMSE) ranging from 0.96 to 0.99 and from 0.28 m to 1.62 m, respectively. The Jason-3 data had the highest correlation with the in situ observation (R² = 0.99) and the lowest correlation (R² = 0.82), with RMSE values ranging from 0.11 m to 1.18 m. With an R² range of 0.93–0.99 and an RMSE range of 0.20–1.05 m, the SARAL/AltiKa mission presented greater accuracy than the Jason altimetry mission. The estimated water levels can be utilized in remote, inaccessible, or ungauged areas and in international transboundary rivers for water storage and river discharge estimations. However, the accuracy of remotely sensed data depends on such factors as along-track distance, water body area, and geographical and terrain conditions near water bodies.     

Enhancement of a global lake and reservoir database to aid climate studies and resource monitoring utilizing satellite radar altimetry

Here, we describe the expansion and enhancement of a large (surface area ≥100 km²) lake and reservoir database (1.GREALD). These efforts have also resulted in the spin-off of two additional databases, one containing lakes and reservoirs (2.GREALD, area 10–99 km²), and the other containing ephemeral lakes (3.GELD, area ≥100 km²). These databases are unique sources for projects that utilize satellite radar altimeter data to monitor surface water levels. While 1.GREALD aims to be a complete catalog, 2.GREALD focuses on reservoirs in response to applied sciences programs that monitor water and energy resources. The creation of 3.GELD has climate change objectives as well as water resources and ecosystem conservation applications. The recording of information pertaining to the potential overpasses (waterbody crossings) of the current and archive satellite altimeters is a primary objective as is the need to highlight any form of controlled water level variation. The permanent water databases now contain 6282 entries, half experience some form of anthropogenic influence and ～430 have been identified as potential climatically sensitive terminal lakes. The revised integral surface area distribution is a power law with exponent ?1.016. Statistics reveal that with altimetric repeat visit times of 10-day to monthly, at least 80% of the permanent water bodies (≥10 km²) have been overflown at some period since the 1990s. Current information on water use and reservoir formation date show that the primary use of the reservoir class is hydroelectric power, and that China, Brazil, India, Turkey, and Vietnam dominate the dam building in recent decades.     

汾河二库坝前垂向表层水温变化规律研究

为探究水库垂向表层水温在年、季、日内的变化特征,给库区水体理化性质分析及养殖业发展提供重要参考,以汾河二库为研究对象,于2016年9月至2017年8月在汾河二库使用10 m长的柔性温度链进行自动化表层水温原型监测,分析了表层水温变化特性。结果表明:汾河二库垂向表层水温分布呈现出明显的季节性变化规律,符合分层型水库表温层特征;秋冬季表层水体水温均匀分布,春夏季水面下1~2 m层存在水温突然升高的现象;通过建立回归模型得知,气温与库面水温之间存在良好的线性相关关系;全年整体库面水温最大值出现时间与气温最大值出现时间一致;不同季节气温对表层水温的影响深度不同,气温升高,水温日变幅值越大,影响深度越大。     

呼伦湖水生态环境遥感监测及其对气候变化的响应

以位于内蒙古的第一大湖-呼伦湖为研究对象,综合应用RS、GIS等现代分析技术手段,通过解译该区1973、1975、1986～2009年近36a的MSS、TM、ETM遥感影像,实现了呼伦湖湖泊水位、面积的遥感动态监测,并建立了关系模型：研究发现呼伦湖面积在1999年达到了1959年以来的最大水面面积2106．95km2后开始骤然下降到2009年的最低水面面积1813．69km2。以24．44km2／a的速度减少了293．26km2,水面面积比1999年最大面积时减少了近14％。水位高程自1999年的最高水位下降了3．94m,是呼伦湖平均水深（9m）的39．4％。收集、整理了呼伦湖及上游流域4个气象站点、3个水文站点近52a的气象、水文资料。分析了呼伦湖区域尺度内的水热状况变化,发现研究区域温度升高。降水呈波动性减少．并且在1999年后温度和降水的变率增大,是呼伦湖面积减小、水位下降的主要原因。最后．分析了气候变化对呼伦湖生态环境的影响效应。并建立了湖泊水位和降水、气温等气象因子以及径流量之间的关系模型．研究可为寒旱区湖泊水环境管理提供参考。     

Seasonality and physical drivers of deep chlorophyll layers in Lake Superior,with implications for a rapidly warming lake

A deep chlorophyll layer (DCL) is a common feature of many deep, oligotrophic lakes including Lake Superior. Mechanisms generating and maintaining DCLs are variable across lakes, and seasonal patterns and relationships of DCL structure to physical variables are not well described. Using vertical profile data for physical and biological variables from western and central Lake Superior, we described seasonal patterns in DCL structure and other physical and biological parameters and applied linear mixed-effects models to determine how different physical factors (surface temperature, thermocline depth, and 1% photosynthetically active radiation (PAR) depth) affect the depth, thickness, maximum concentration, and integrated chlorophyll of the DCL. We observed clear seasonal patterns in the development and degradation of the DCL that coincide with seasonal changes in light and temperature. Modeling analysis using linear mixed-effects models showed that the DCL thickness was best predicted by surface temperature (R² = 0.51) followed by thermocline depth (R² = 0.36), and the deep chlorophyll maximum (DCM) concentration was best predicted by surface temperature (R² = 0.26). The 1% PAR depth was not implicated as an important predictor, but observations from seasonal data suggest that it plays a role in the depth of the DCM. While no relationship was found between surface temperature and DCL-integrated chlorophyll, DCL thickness decreased and DCM concentration increased with increasing surface temperature, which could have implications for productivity in the DCL as the lake continues to warm.     

Complex summer water temperature behaviour below a UK regulating reservoir

Detailed monitoring over a 15-year period has revealed considerable complexity in summer water temperature behaviour below Wimbleball Lake, a large regulating reservoir in south-west England. Continuous records of water temperature in the regulated River Haddeo and the neighbouring unregulated River Pulham, coupled with information on flow in the tailbay of the dam and on the thermal structure of the impounded water body, reveal that the nature and extent of the thermal modification below the reservoir varies from day to day, month to month and year to year. This complexity largely reflects changes in the balance between a springflow and a direct reservoir release component of the compensation runoff, which arise in response to variations within and between summers in hydrometeorological conditions and in the way the reservoir is operated to regulate downstream flows. Temperature behaviour in the regulated river is made more complicated by progressive warming of the springflow during the summer months, the thermal inertia of the stored water, the development of thermal stratification within the reservoir and the alteration of release depth, although the detailed effect of some of these factors depends on weather conditions, the use of destratification equipment and the rapidity of reservoir drawdown. © 1997 John Wiley & Sons, Ltd.     

Modeling the impacts of climate change on the thermal and oxygen dynamics of Lake Volta

Climatic changes influence the thermal and oxygen dynamics of a lake and thus its ecological functioning. The impacts of climatic changes on tropical lakes are so far poorly studied and the extent of the effects is therefore uncertain, most investigations describing only potential effects. In this study, we applied the one-dimensional lake ecosystem model GOTM-ERGOM to quantify the effects of climate change on thermal stratification, oxygen dynamics, and primary production in meso-oligotrophic Lake Volta. GOTM-ERGOM was calibrated and validated using two years of observed data. The validated model was used to evaluate a series of future climate change scenarios. The model simulations showed good agreement with observed water temperature, dissolved oxygen and chlorophyll-a and indicated intensified stratification and reduced oxygen levels in the productive water layers of the lake. However, the longer-lasting stratification (prolonged stability) did not translate into permanent stratification. A relatively small (1?m) upward shift of thermocline depth resulted in an 8%–12% volume loss of the oxygen-rich upper mixed layer, which may be significant for the fisheries of the lake as it diminishes the size of suitable fish habitats. Light limitation of primary production renders the lake somewhat resilient to intensive algae blooms, as traceable in both the present and in the future climate scenarios. In the long term, the ongoing climate change may affect riparian communities that depend on the lake's fisheries for their livelihood. In consequence, future lake management strategies for implementation need to account for the impacts of future climate change.     

Summer water circulation in Frenchman's Bay,a shallow coastal embayment connected to Lake Ontario

Water movements due to temperature gradients and short-term water level fluctuations control the flushing timescale of the many shallow embayments in the Great Lakes. In this article the water circulation within Frenchman's Bay is reported and estimates made of the hydraulic flushing timescale. This shallow freshwater embayment is permanently connected to Lake Ontario through a channel of width 30 m and depth of 2 m. The water flushing timescale is estimated by using a salt mass budget, leading to a flushing timescale of 7–10 days. During the summer, the exchange of water between the bay and the lake can be driven by a combination of horizontal thermal gradients, and by small but ubiquitous 1–5 cm oscillations in the water level of Lake Ontario. The water movements that are predicted due to water level fluctuations (caused by seiches, storm surges and tides) and due to horizontal thermal gradients leads to estimates of the flushing time of water within Frenchman's Bay in the range of 12–13.5 days, i.e. the same order of magnitude as the salt budget. One consequence of the temperature driven exchange flows is to cause strong and persistent temperature stratification within the bay, which in combination with high nutrient loading and low winds in summer leads to frequent anoxic events.     

THE SOLAR‐TO‐STREAM POWER RATIO: A DIMENSIONLESS NUMBER EXPLAINING DIEL FLUCTUATIONS OF TEMPERATURE IN MESOSCALE RIVERS

The diel variation of temperature in mesoscale river reaches (catchment area > 1000 km²) is analysed using concurrent measurements of water temperature and of those meteorological (incident short‐wave radiation, air temperature, relative humidity and wind speed variables) and hydraulic variables (streamflow, top width, channel slope and flow depth) controlling the thermal regime. Measurements were taken along two river reaches located in central Chile, on the Itata (11 290 km², Strahler's order 6, reach length 30 km, Q_bankfull = 400 m³ s^?1) and Vergara (4340 km², Strahler's order 5, reach length 20 km, Q_bankfull = 85 m³ s^?1) rivers. The measuring frequency was 15 min. The relevant energy fluxes at the air–water interface, that is, atmospheric long‐wave radiation, net short‐wave radiation, radiation emitted by the water body, evaporation (latent heat) and conduction heat are computed and analysed for four scenarios of 12 days duration each, representing typical conditions for the austral winter, spring, summer and autumn. We find large differences in the diel river temperature range between the two sites and across seasons (and thus, flows and meteorological conditions), as reported in previous studies, but no clear relationship with the controlling variables is overtly observed. Following a dimensional analysis, we obtain a dimensionless parameter corresponding to the ratio of solar‐to‐stream power, which adequately explains the diel variation of water temperature in mesoscale rivers. A number of our own measurements as well as literature data are used for preliminary testing of the proposed parameter. This easy‐to‐compute number is shown to predict quite well all of the cases, constituting a simple and useful criterion to estimate a priori the magnitude of temperature diel variations in a river reach, given prevailing meteorological (daily maximum solar radiation) and hydrologic–hydraulic (streamflow, mean top width) conditions. Copyright © 2012 John Wiley & Sons, Ltd.     

Initial insights on the thermal ecology of lake whitefish in northwestern Lake Michigan

Lake whitefish Coregonus clupeaformis are a native coldwater species supporting important recreational and commercial fisheries in the Laurentian Great Lakes. Climate-related changes in water temperature may have important implications for the future sustainability of these fisheries. However, projecting future habitat availability is difficult because limited information is available on lake whitefish thermal ecology in the region. In this study, archival temperature loggers were implanted into 400 lake whitefish from northwestern Lake Michigan, including Green Bay, during October–November 2017. Loggers recorded temperature for 11 months at 4-hr intervals. Thirteen recovered temperature loggers were used in analyses. In winter (1 December–31 March), temperatures occupied by lake whitefish ranged from 0 to 8.0 °C, while in spring (1 April–31 May) temperatures ranged from 0 to 20.0 °C. In summer (1 June–15 September) and fall (16 September–7 November), lake whitefish occupied temperatures of 4–21.5 and 4–21.0 °C, respectively. Average temperatures in summer (10.8 °C) were within the previously proposed optimal temperature range (10–14 °C) and broad thermal niche (7–17 °C); however, 58% of observations were outside the optimal temperature range and 11% of observations were outside the broad thermal niche. Our results suggest that lake whitefish from northwestern Lake Michigan inhabit temperatures both above and below previously reported expected temperature ranges. This study provides initial insights on lake whitefish thermal ecology in Lake Michigan and can be used as a baseline for future work aimed at determining how lake whitefish habitat availability may change in the future.     

Seasonal variation in light penetration and subsurface chlorophyll-α in southern Lake Michigan observed by a glider

The Cooperative Institute for Great Lakes Research (CIGLR) in collaboration with the Great Lakes Observing System and National Oceanic and Atmospheric Administration, Great Lakes Environmental Research Laboratory (NOAA GLERL) deployed an autonomous underwater glider in southern Lake Michigan several times per year between 2012 and 2019 to collect offshore (>30 m depth) limnological measurements, including temperature, photosynthetically active radiation (beginning during 2015), and chlorophyll-α fluorescence. From these data, we calculated mixed layer depth, several measures of light penetration (diffuse attenuation coefficient, first optical depth, euphotic zone depth), and depth of the subsurface chlorophyll-α maxima. During summer, mean offshore mixed layer depth was typically 10–15 m, K_d for PAR was 0.1–0.17 m^?1, first optical depth was 6–9 m, euphotic zone depth was 35–40 m, and depth of subsurface chlorophyll-α maxima was 30–35 m. We also observed substantial spatial and temporal variation in these values across the basin and within and among seasons. Glider-based observations provide a wider horizontal and vertical perspective than other methods (e.g., ship- and satellite-based observations, buoys, and fixed moorings), and are therefore a valuable, complementary tool for Great Lakes limnology. The set of observations reported here provide seasonal and basin-scale information that may help to identify anomalies useful for future glider-assisted investigation into the role of biophysical processes in Great Lakes limnology and ecology.     

Optical and biological properties of Lake Ülemiste, a water reservoir of the city of Tallinn I: Water transparency and optically active substances in the water

Optical and biological measurements were performed in Lake Ülemiste in the summer of 1997 (four measurement days), and from May to October in 1998 (12 measurement days). This kind of data, describing the type and amount of optically active substances in the water, phytoplankton characteristics, the underwater light field, and temperature–oxygen situation in the lake are necessary when estimating the ecological state of the lake. Lake Ülemiste is the main drinking water reservoir of Tallinn, the capital of Estonia. Phytoplankton abundance and biomass, chlorophyll a and suspended matter were determined from collected Lake Ülemiste water samples in the laboratory. Spectrophotometrical processing of the filtered and unfiltered water was carried out to describe the beam attenuation coefficient spectra and optical influence of yellow substance in the water. Vertical profiles of downwelling irradiance of the PAR (400–700 m) region of the spectrum (and from these data the averaged over depth diffuse attenuation coefficient) were determined. The relative transparency of the water was estimated by using a Secchi disk. Passive optical remote measurements were episodically made from aboard a boat. Results obtained confirmed that Lake Ülemiste is turbid (almost hypertrophic), comparable with most turbid lakes in Estonia (e.g. Lake Võrtsjärv) and Finland (Lake Tuusulanjärvi). Its chlorophyll a content varied within the range 13–121 mg m^–3, phytoplankton biomass 3–107 mg L^–1, phytoplankton abundance 65 000–999 000 mL^–1, suspended matter 8–34 mg L^–1, effective concentration of the yellow substance 6–30 mg L^–1, diffuse attenuation coefficient of light in the PAR region 1.0–3.3 m^–1 and a Secchi disk depth of 0.5–1.75 m. The temporal variation of the spatial averages of these parameters during 1998 was analysed. Almost all characteristics showed an increase from May to midsummer with a maximum in late July or in August (correspondingly the Secchi depth values were minimal in late summer). The amount of yellow substance was an exception, which was nearly constant during the observation period. The maximum level of chlorophyll a content in July and August 1998 markedly exceeded that in 1997, despite the fact that the summer of 1997 was sunny, but the summer of 1998 was cold and rainy. The correlative relationships between the different parameters were investigated together with the respective data for other lakes. They show that the data of Lake Ülemiste supplemented the correlation graph in the region of turbid lakes, whereby in all cases the correlation coefficient increased following the addition of Lake Ülemiste data. The highest correlation coefficient was obtained when light attenuation coefficient values were correlated with a sum of weighted concentrations of chlorophyll a, yellow substance and suspended matter (multiple regression analysis).     

官厅水库夏季叶绿素a浓度短时分布特征

分析官厅水库夏季(8月份)叶绿素a浓度短时变化特征,对于水库水体富营养化研究具有重要意义。利用以分钟为步长的水温、水深及叶绿素a浓度实测数据及气象数据,对叶绿素a浓度垂向及昼夜变化特征进行了分析。结果表明:叶绿素a浓度最高值是表层浓度的1.5~2倍,其出现位置与温度跃层位置基本一致。叶绿素a浓度具有昼夜周期性变化特征。白昼,温度跃层以上叶绿素a浓度随水深增加逐渐增加,在温度跃层以下,随水深增加逐渐降低;夜间,温跃层以上叶绿素a浓度基本稳定不变,在温度跃层以下与白昼变化趋势相同。表层水体叶绿素a浓度夜间大于白天,且与光照及表层水温呈负相关。水库叶绿素a浓度短时分布特征主要与温度分层以及限制性营养盐磷的释放有关。     

A floodplain forest mosquito community after man-made moisture changes (Culicidae,Diptera)

This paper summarizes data on the response of mosquitoes to changed water regime following extensive water management alterations on the southern Moravian Dyje and Morava rivers. Maximum groundwater level dropped by an average of 90 cm, previous regular floods were prevented, and the area covered by floodplain forests was significantly decreased due to the construction of three reservoirs. The original mosquito community of temporary water bodies (Aedes) is in retreat both quantitatively and, in places, qualitatively. Of the initial number of six species causing plagues, only Ae. vexans has remained (63.3 per cent), the rest occurring at percentages of only 0.1–4.1 per cent. After flood regulation, the breeding grounds of the species developing in temporary water bodies has filled only during periods of sufficiently high groundwater levels and sufficient saturation of soil with rainfall. A prospective group in this area includes mosquito species developing in permanent water bodies (Culex, Anopheles, Culiseta, etc.). This group is increasing in numbers and species richness. After the completion of three reservoirs, Anopheles spp. spread throughout the study area. They develop in the waters accompanying the reservoirs, especially in channels rich in vegetation, in which they attain densities of up to 39.9 larvae per 100 cm² water surface. The drier environment, and drier and warmer climate prevailing in recent years have probably influenced the occurrence of southern species in the area, notably Culex martinii and Uranotaenia unguiculata. Long-term quantitative collections of mosquitoes enable first note to be made on the possible fluctuation mechanisms, previously obscured by the essential influence of floods on community development.     

Can water level stations be used for thermal assessment in aquatic ecosystem?

Many studies focus on stream water temperature (WT) because it is considered a key ecological factor. However, few of them have investigated the use of WT data from water level monitoring networks, which often measure WT as ancillary data. Our study was conducted in southern Belgium at a high temporal resolution with continuous data recorded at intervals of 10 min between 2012 and 2016 and large spatial scale greater than 16,000 km². This study aimed to assess whether a regional water level network (140 stations) is reliable for continuous WT monitoring based on a Bland–Altman analysis with WT collected through a European monitoring network (Water Framework Directive). This study also investigates whether WT data acquired by water level stations can be used to perform both state‐of‐the‐art visualization of thermal regimes and spatio‐temporal queries for specific ecological monitoring. We found that the water level stations were reliable tools in recording continuous WT in the streams of the study area. The temperature difference between the two WT monitoring networks was ?0.57°C on average. Our positive results promote the use of WT from water level stations in order to globally characterize the thermal regime of streams as well as to provide spatial or temporal information on this regime at high frequencies. As an example, our data showed the effectiveness for brown trout (Salmo trutta fario L.) in spatializing thermal risk areas related to the thermal requirement of this fish species; in 2015, 19% of stations located in brown trout fish zone recorded temperatures above 25°C.     

A study of physical parameters at the spawning sites of the european grayling (Thymallus thymallus L.)

An investigation of the spawning sites of European grayling, Thymallus thymallus (L.), in two Swedish rivers revealed narrow range in their physical parameters. The bottom substrate was composed of 10–20 per cent sand, 50–70 per cent gravel (<2cm), 20–30 per cent stones (2–10cm), and a few bigger stones (>10cm in diameter). Eggs were found only where the gravel thickness was more than 5 cm. The depth varied between 30–50cm. Mean depth for 22 different spawning sites was 36cm. The water velocity varied between 23–90cm s^?1, average 54 cm s^?1. The temperature at the start of spawning was 3.9°C and at the end it was about 9°C. The results indicate the narrow range in physical parameters at the spawning sites. Small changes in the environment caused by regulation of river flow or water level may disturb the spawning behaviour of grayling and cause a decrease in population size or even extinction.