At‐A‐Station Hydraulic Geometry Simulator

Presented in this paper is a hydraulic model that combines a rational regime theory with an at‐a‐station hydraulic geometry simulator (ASHGS) to predict reach‐averaged hydraulic conditions for flows up to but not exceeding the bankfull stage. The hydraulic conditions determined by ASHGS can be paired with an empirical joint frequency distribution equation and applicable habitat suitability indices to generate weighted usable area (WUA) as a function of flow. ASHGS was tested against a 2‐dimensional hydrodynamic model (River2D) of a mid‐size channel in the Interior Region of British Columbia. By linking ASHGS to a regime model, it becomes possible to evaluate the direction and magnitude of habitat changes associated with a wide range of environmental changes. Our regime model considers flow regime, sediment supply, and riparian vegetation: these governing variables can be used to simulate responses to forest fire, flow regulation and changes in climate and land use. Practitioners can examine ‘what‐if’ scenarios that otherwise would be too expensive and time consuming to fully explore. The model boundaries of commonly used data‐intensive hydraulic habitat models (e.g. PHABSIM) are not easily adjusted and such models are not designed to estimate future morphological and hydraulic habitat conditions in rivers the undergo significant channel restructuring. The proposed model has the potential to become an accepted flow assessment tool amongst practitioners due to modest data requirements, user‐friendliness, and large spatial applicability; it can be used to conduct preliminary assessments of channel altering projects and determine if in‐depth habitat assessments are justified.     

GRADIENTS IN THE BIOPHYSICAL STRUCTURE OF URBAN RIVERS AND THEIR ASSOCIATION WITH RIVER CHANNEL ENGINEERING

Urban rivers are often engineered to increase flood conveyance and stabilize channel size and position. This paper analyses habitat surveys of 180 urban river stretches of differing engineering type from four river basins (river Tame, West Midlands, UK; tributaries of the lower river Thames, UK; river Botic, Prague, Czech Republic; river Emscher, North‐Rhine Westphalia, Germany). Kruskal–Wallis tests identify significant differences in extent and/or frequency of flow types, bank and bed physical habitats, and vegetation characteristics associated with different styles of engineering. Principal Components Analysis identifies four key environmental gradients in the data set: sediment supply and retention; extent and diversity of in‐channel vegetation and riparian trees; bed and bank sediment calibre; flow type energy and complexity. These gradients discriminate stretches of differing planform, cross section and reinforcement and are significantly correlated with indices of degree and type of bank and bed reinforcement, pollution and presence of alien nuisance plant species. The analytical results illustrate statistically significant associations between different styles and levels of engineering intervention and the number and nature of physical habitats present in urban rivers. The results provide a basis for filtering sites for potential remedial measures prior to site‐specific surveys and modelling, for comparing sites and for tracking trajectories of change at sites that are subject to changes in channel engineering. They provide evidence that river condition and degree of engineering are not inversely related in a simple linear way, and that engineering of urban river channels, in the form of mixed, patchy reinforcement can contribute a great deal to habitat diversity where other controls on flow heterogeneity are more difficult to manipulate. Copyright © 2011 John Wiley & Sons, Ltd.     

Habitat suitability curves for brown trout (Salmo trutta fario L.) in the River Adda,Northern Italy: comparing univariate and multivariate approaches

Habitat suitability of brown trout (Salmo trutta fario) was studied in the upper portion of the Adda River, Northern Italy. Measurements were made for 528 individuals distributed in two life‐stage classes, adult and juvenile, based on body length. In order to provide basic biological information for the physical habitat simulation (PHABSIM) system of the instream flow incremental methodology (IFIM) in the Italian regulated rivers, habitat suitability curves (HSCs) have been developed with respect to several microhabitat riverine parameters. Initially, current velocity, water depth, substrate class size and cover were analysed with an univariate approach, then bivariate habitat suitability models were developed from depth and velocity data. The comparison of experimental univariate HSCs with those from the literature outlined some differences that can essentially be explained by characteristics of the investigated river, confirming the necessity of using site‐specific curves in relation to each experimental study area. To compare the univariate and bivariate approaches, the weighted usable area (WUA)–discharge relationships were calculated using both types of HSCs. Response curves obtained from the two approaches turned out to be quite different. In PHABSIM habitat modelling, HSCs univariate functions need to be aggregated to produce the WUA–discharge relationship. A multiplicative criterion is generally used for the combined suitability factor; by means of this aggregation criterion all variables have equal weight. According to bivariate models, depth is much more important than velocity in defining habitat suitability requirements. Copyright © 2001 John Wiley & Sons, Ltd.     

Linking a spatially explicit watershed model (SWAT) with an in‐stream fish habitat model (PHABSIM): A case study of setting minimum flows and levels in a low gradient,sub‐tropical river

As changes in landuse and the demand for water accelerate, regulators and resource managers are increasingly asked to evaluate water allocation against the need for protection of in‐stream habitat. In the United States, only a small number of river basins have the long‐term hydrograph data needed to make these assessments. This paper presents an example of how to bridge the conceptual and physical divide between GIS‐based watershed modelling of basin‐discharge and in‐stream hydraulic habitat models. Specifically, we used a Soil and Water Assessment Tool (SWAT) model for the Hillsborough River to produce data for use in a Physical HABitat SIMulation (PHABSIM) model of the same river. This coupling of models allowed us to develop long‐term discharge data in ungauged river systems based on watershed characteristics and precipitation records. However this approach is not without important limitations. Results confirm that accuracy of the SWAT‐predicted hydrograph declines significantly when either the DEM resolution becomes too coarse or if DEM data are resampled to a coarser or finer resolution. This is due to both changes in the size and shape of the river basin with the varying DEMs and subsequent shifts in the proportions of land use, soils and elevation. Results show the use of 30 m DEMs produced hydrographic patterns amenable for using in‐stream habitat protocols like PHABSIM model, especially where little or no hydrographic and land use information exists. Copyright © 2010 John Wiley & Sons, Ltd.     

Predicted Native Fish Response,Potential Rewards and Risks from Flow Alteration in a New Mexico Arid Mountain Stream

Water and natural resource managers are concerned with evaluating how fish habitat and populations may respond to water diversions and small‐scale flow augmentations. We used two‐dimensional hydraulic models, habitat suitability curves and an individual‐based population viability model to assess whether flow augmentations of about 0.28–0.57 m³/s would create suitable habitat for federally listed native fish loach minnow Rhinichthys cobitis and spikedace Meda fulgida in a reach of the Gila River, New Mexico, and then examined how fish population viability may change under a variety of colonization and extinction scenarios. These simulations help to inform water management decisions in a reach of the Gila River where river diversions currently exist and new diversions and augmentations are being proposed. Our results suggest that the flow augmentations evaluated will result in small changes (on average across life stages, ?0.22% to 4.06%) in suitable habitat for loach minnow and spikedace depending on augmentation scenario and fish life stage. While these percent changes are small, they would result in a reduction in the dewatering of the river channel in a river reach where native fish abundance is thought to be low. Actual native fish responses to these habitat changes are unknown; however, these flow augmentations could potentially allow these native species to re‐colonize this river segment from upstream or downstream sources increasing species distribution and likely population viability. Maintaining viable populations of native fish in this river reach is dependent on complex factors including persistence of suitable habitat for multiple life stages, connectivity with other populations and minimizing risk of invasion from non‐native species. We recommend that these predictions from the habitat and population models be tested and verified in an adaptive management framework linking modelling, experimental management, monitoring and reassessment to inform water management decisions in the Gila River. Copyright © 2017 John Wiley & Sons, Ltd.     

Comparison of MesoHABSIM with two microhabitat models (PHABSIM and HARPHA)

This study demonstrates how using different habitat models can influence the results of instream habitat assessment and conclusions for river management. We used three models for a portion of the Quinebaug River (Connecticut and Massachusetts, USA): a simplistic microhabitat model with univariate habitat‐use criteria and substrate‐based channel index (the Physical Habitat Simulation Model (PHABSIM)); a microhabitat model using multivariate criteria including a wide range of cover attributes (HARPHA); and a mesohabitat model with multivariate habitat‐suitability criteria (MesoHABSIM). The flow‐habitat rating curves produced by each model were compared at two scales: site and study segment. To investigate the impact of model choice on answering questions such as which location or flow provides more habitat, we applied Spearman's correlation of ranks. The relationship between habitat‐suitability predictions and fish presence at the same location was investigated with dedicated fish observations. The study showed that: (1) of the tested models, only MesoHABSIM predictions correlated with fish observations; (2) the variation within microscale models (PHABSIM and HARPHA) was greater than between micro‐and mesoscale models (HARPHA and MesoHABSIM); and (3) simple univariate habitat‐use criteria provided the largest source of discrepancies among the models. We suggest that these differences may lead to erroneous conclusions, especially if flow‐habitat rating curve analysis is considered an endpoint of instream flow study. Copyright © 2007 John Wiley & Sons, Ltd.     

THE INFLUENCE OF VARIABLE HABITAT SUITABILITY CRITERIA ON PHABSIM HABITAT INDEX RESULTS

The Physical Habitat Simulation System (PHABSIM) still probably remains as the most widespread habitat method used to establish inflow standards or to link habitat temporal variations with fish population dynamics. However, statistical uncertainties around the PHABSIM main output, the weighted usable area (WUA) over discharge curves, are usually ignored. Here, we assess the uncertainty in WUA curves and derived habitat duration curves induced by the variability around the PHABSIM biological model, the habitat suitability criteria, using brown trout Salmo trutta as the model species. Bootstrap analyses showed that the uncertainty around the WUA curves was rather high when bootstrap sample (BS) size was low and differed among age classes, being generally lower for young‐of‐the‐year (YOY). Width of 95% confidence intervals for maximum WUA magnitude increased with decreasing BS size, ranging from 19.3% for YOY trout at the largest BS size (40 transects, 270 habitat use observations) to 146% for juveniles at the smallest BS size (nine transects, 60 habitat use observations). The uncertainty arose primarily from the construction of the channel index variable. Nevertheless, results showed that the uncertainty in WUA values could be reduced down to acceptable levels by using general functional channel index categories. Likewise, the shape of WUA curves was also highly variable when BS was small. These patterns resulted in habitat duration curves being highly uncertain, much more in their amplitude than in their shape. Uncertainty about the flows corresponding to different habitat exceedance values increased with decreasing probability of exceedance. Width of peak flow confidence intervals ranged from 3.3% for YOY trout at the largest BS size to 226% for adults at the smallest BS size. Yet such levels of uncertainty do not necessarily entail critical errors in the decision‐making process because large variability in flow peak does not necessarily lead to large variability in WUA magnitude.     

Increasing the availability and spatial variation of spawning habitats through ascending baseflows

Precipitation in fall and winter is important to recharge aquifers in Northern California and the Pacific Northwestern United States, causing the baseflow in rivers ascend during the time when Chinook salmon (Oncorhynchus tshawytscha) construct redds. Herein, we evaluate the availability of spawning habitats under a constant streamflow common in regulated rivers against ascending baseflows patterned from free‐flowing rivers. A binomial logistic regression model was applied to predict the suitability of redd locations based on physical characteristics. Next, two‐dimensional hydrodynamic habitat models were developed at two locations representing a broad range of channel forms common in large rivers. Hydrodynamic and habitat models were leveraged together to simulate the quality, amount, and spatial distribution of spawning habitat at a series of individual flow rates, as well as the combined effect of those flow rates through a spawning season with ascending baseflows. Ascending baseflows increased the abundance of spawning habitat over individual streamflows at a site where the river channel is confined by levee‐like features. However, improvements were greater at an unconfined site that facilitated lateral connectivity and greater expansion of wetted channel area as streamflows increased. Ascending baseflows provided spatial separation in preferred habitats over a spawning season, which may reduce the risk of superimposition among runs or among species. Ascending baseflows provided a benefit across the range of hydrologic regimes in a 100‐year gauge record ranging from 20% to 122% improvements in habitat area over low streamflows that are currently used to manage for spawning habitat. Although replicating natural flow regimes in managed systems can be impossible or impractical, these results demonstrate that incorporating elements of the natural flow regime like ascending baseflows can benefit the restoration and conservation of riverine species.     

Interactive effects of cover and hydraulics on brown trout habitat selection patterns

Habitat modelling results are extremely sensitive to the habitat suitability criteria (HSC) used in the simulations. HSCs are usually expressed as univariate habitat suitability curves, although such univariate approach has been long questioned, since overlooking interactions between hydraulic variables may misrepresent the complexity of fish behaviour in habitat selection. It could lead to adopt erroneous flow management decisions based on misleading results. Furthermore, the interactive effects of hydraulic variables on habitat selection may be driven by the structural features of the channel, which determine cover availability. Therefore, we compared brown trout habitat selection patterns through multivariate resource selection functions (RSFs) in structurally contrasting rivers to unveil the interactive effects of hydraulics and cover elements and their consequences in univariate HSC results. Microhabitat preferences of young‐of‐the‐year (0+) trout were similar across fast and slow waters, meanwhile juvenile (1+) and adult (>1+) preferences significantly changed. RSFs for young‐of‐the‐year trout were consistent with univariate results and did not differ among water types. However, RSFs for older trout varied among water types and revealed complex interactions among hydraulic variables and between hydraulics and structural elements, which were not described accurately by univariate curves. Therefore, results suggest that interactions between water depth and current velocity have a significant effect on habitat selection patterns in juvenile and adult brown trout, this effect being controlled by cover availability. Copyright © 2008 John Wiley & Sons, Ltd.     

Evaluating Uncertainty in Physical Habitat Modelling in a High‐Gradient Mountain Stream

Predictions of habitat‐based assessment methods that are used to determine instream flow requirements for aquatic biota are uncertain, but instream flow practitioners and managers often ignore those uncertainties. Two commonly recognized uncertainties arise from (i) estimating the way in which physical habitat within a river changes with discharge and (ii) the suitability of certain types of physical habitat for organisms. We explored how these sources of uncertainty affect confidence in the results of the British Columbia Instream Flow Methodology (BCIFM), which is a commonly used transect‐based habitat assessment tool for small‐scale water diversions. We calculated the chance of different magnitudes of habitat loss resulting from water diversion using a high‐gradient reach of the North Alouette River, BC, as a case study. We found that uncertainty in habitat suitability indices for juvenile rainbow trout generally dominated uncertainty in the results of the BCIFM when large (>15) numbers of transects were used. In contrast, with small numbers of transects, variation in physical habitat among sampled transects was the major source of uncertainty in the results of the BCIFM. Presentations of results of the BCIFM in terms of probabilities of different amounts of habitat loss for a given flow can help managers prescribe instream flow requirements based on their risk tolerance for fish habitat loss. Copyright © 2015 John Wiley & Sons, Ltd.     

Effect of flow‐related substrate alteration on physical habitat: a case study of the endemic river loach Sinogastromyzon puliensis (Cypriniformes,Homalopteridae) downstream of Chi‐Chi Diversion Weir,Chou‐Shui Creek,Taiwan

This paper addresses the variation of substrate status with flow in the framework of habitat simulation. The Physical Habitat Simulation System (PHABSIM) was developed as a tool for water management and has become one of the most popular methods worldwide. Despite its many advantages, the variation of channel properties with flow is not addressed in PHABSIM. We modified PHABSIM by incorporating a sub‐program that can perform substrate analyses under five different schemes for evaluating the substrate suitability. These schemes include the conventional methods to specify the substrate attribute (scheme 1) and to use the mean grain size of the original bed material for determining the substrate suitability (scheme 2), or employ the threshold conditions for sediment entrainment to evaluate the nominal grain size of the stable substrate (schemes 3–5). As a case study, Sinogastromyzon puliensis in Chou‐Shui Creek (Taiwan) is selected as the target species for the habitat simulation. For schemes 1 and 2, the substrate attribute specified for each cell of a cross‐section does not change with flow. The nominal grain size of the stable armour material (scheme 5) is generally greater than the nominal grain sizes of the above‐critical bed material (scheme 3) and the static armour layer (scheme 4). The simulation results indicate that the suitability values specified in scheme 1 and the resulting habitat area are on the optimistic side, whereas scheme 2 tends to undervalue the suitability of the hydraulically worked substrate. The overall variation trends of the weighted usable area (or percentage usable area) curves for different substrate schemes are similar, especially in the in‐phase regions of the water velocity and depth. The results of this study appear to imply that the flow‐related habitat analyses are not usually sensitive to substrate. Copyright © 2002 John Wiley & Sons, Ltd.     

A probability-based model to quantify the impact of hydropeaking on habitat suitability in rivers

A negative effect of hydropower on river environment includes rapid changes in flow and habitat conditions. Any sudden flow change could force fish to move towards a refuge area in a short period of time, causing serious disturbances in the life cycle of the fish. A probability-based model was developed to quantify the impact of hydropeaking on habitat suitability for two fish species, brown trout (Salamo trutta) and Grayling (Thymallus thymallus). The model used habitat preference curves, river velocity and depth to develop the suitability maps. The suitability maps reveal that habitat suitability deteriorates as flow increases in the studied part of the river. The probability model showed that, on average, suitability indices are higher for adult grayling than juvenile trout in hydropeaking events in this part of the river. The method developed shows the potential to be used in river management and the evaluation of hydropeaking impacts in river systems affected by hydropower.     

Exploring functional flow heterogeneity in regulated flow regime: Fish species turnover along hydraulic gradients in an artificial waterway network

Humans have altered river flows leading to a loss of connection with floodplain habitats. The expansion of agriculture in floodplains has resulted in landscapes dominated by irrigated farmland. A key challenge in water management is to conserve existing ecological communities and habitat heterogeneity, while simultaneously maintaining engineered infrastructure for agriculture. In this study, we focused on an artificial channel network for irrigation with a regulated flow regime and its function as habitat for various fish species. Differences in hydraulic conditions among channels and compositional changes in fish species were examined to clarify functional flow heterogeneity. Analyzed using pairwise Simpson dissimilarity among sampling reaches, species turnover was positively associated with Froude number (flow intensity) differences at intermediate discharges, and with differences in cross-sectional areas (flow magnitude) at low discharges. Drastic changes in inflows should be considered for the effective conservation of flow heterogeneity, even under a regulated flow regime. Improved engineering design to manage the hydraulic environment is one option for maintaining the ecological value of lateral waterbodies in human-dominated landscapes. Our findings provide insights into the importance of functional flow heterogeneity to conserve fish species diversity.     

Low‐Head Dam Impacts on Habitat and the Functional Composition of Fish Communities

The natural flow regime of many rivers in the USA has been impacted by anthropogenic structures. This loss of connectivity plays a role in shaping river ecosystems by altering physical habitat characteristics and shaping fish assemblages. Although the impacts of large dams on river systems are well documented, studies on the effects of low‐head dams using a functional guild approach have been fewer. We assessed river habitat quality and fish community structure at 12 sites on two rivers; the study sites included two sites below each dam, two sites in the pool above each dam and two sites upstream of the pool extent. Fish communities were sampled from 2012 to 2015 using a multi‐gear approach in spring and fall seasons. We aggregated fishes into habitat and reproductive guilds in order to ascertain dams' effects on groups of fishes that respond similarly to environmental variation. We found that habitat quality was significantly poorer in the artificial pools created above the dams than all other sampling sites. Fast riffle specialist taxa were most abundant in high‐quality riffle habitats farthest from the dams, while fast generalists and pelagophils were largely restricted to areas below the downstream‐most impoundment. Overall, these dams play a substantial role in shaping habitat, which impacts fish community composition on a functional level. Utilizing this functional approach enables us to mechanistically link the effects of impoundments to the structure of fish communities and form generalizations that can be applied to other systems. Copyright © 2017 John Wiley & Sons, Ltd.     

PHABSIM modelling of Atlantic salmon spawning habitat in an upland stream: testing the influence of habitat suitability indices on model output

This paper reports the findings of research designed to assess the ability of PHABSIM to predict Atlantic salmon Salmo salar spawning habitat in the Girnock Burn, a tributary of the River Dee in northeast Scotland. It used an 18‐year spawning data record to assess: (a) the ability of PHABSIM to predict between‐year differences in the availability of habitat at the study site; (b) the ability of PHABSIM to predict patterns of relative suitability across the site; and (c) the influence of different Habitat Suitability Indices (HSIs) on the model's predictions with respect to (a) and (b). Predictions of between‐year and within‐site habitat availability based on ‘utilization’ and ‘preference’ HSIs developed in the Dee catchment corresponded significantly (chi‐squared and regression tests, P < 0.05) with the use of the site by spawning fish. However, predictions based on utilization HSIs developed in streams in southern England did not correspond significantly with patterns of site use. Results of the study indicate that PHABSIM is capable of predicting Atlantic salmon spawning habitat in upland streams such as the Girnock, but that the use of appropriate HSIs is critical. Copyright © 2005 John Wiley & Sons, Ltd.     

Using habitat guilds to develop habitat suitability criteria for a warmwater stream fish assemblage

The diversity of fish species found in warmwater stream systems provides a perplexing challenge when selecting species for assessment of instream flow needs from physical habitat analyses. In this paper we examined the feasibility of developing habitat suitability criteria (HSC) for the entire fish community of a warmwater stream using habitat guilds. Each species was placed a priori into a guild structure and habitat data were collected for depth, velocity, Froude number, distance to cover, embeddedness and dominant and subdominant substrate. Correct guild classification was tested with linear discriminant analysis for each species. Correct classification based on habitat‐use data was highest for riffle and pool‐cover guilds, whereas the fast‐generalist and pool‐run classes, the broader niche guilds, were more frequently misclassified. Variables most important for discriminating guilds were Froude number, velocity and depth in that order. Nonparametric tolerance limits were used to develop guild suitability criteria for continuous variables and the Strauss linear index was used for categorical variables. We recommend the use of a wide array of variables to establish more accurate habitat analysis. Additionally, guild HSC can be developed with similar effort to that needed to develop HSC for a small number of individual species. Results indicate that a habitat guild structure can be successfully transferred to another river basin and that habitats for a diverse fish assemblage can be adequately described by a small number of habitat guilds. This approach represents an alternative for incorporating entire fish assemblages into habitat analyses of warmwater stream systems. Copyright © 2010 John Wiley & Sons, Ltd.     

Low-head dam removal increases functional diversity of stream fish assemblages

Despite the growing number of dam removals, few have been studied to understand their impacts on stream fish communities. An even smaller proportion of dam removal studies focus on the impacts of low-head dam removals, although they are the most common type of dam. Instead, the majority of removal studies focus on the impacts of larger dams. In this study, two previously impounded Illinois Rivers were monitored to assess the impacts of low-head dam removal on the functional assemblage of stream fishes. Study sites were sampled each fall from 2012–2015 (pre-dam removal) and 2018–2020 (post-dam removal) in three locations: the tailrace, impoundment, and river channel. Fishes were aggregated into habitat and reproductive guilds, relating community changes to habitat, environmental metrics, and stream quality. Prior to removal, the slackwater guild was the most prevalent habitat guild throughout both rivers, while nest builders and benthic spawners were the most abundant reproductive guilds. During the two years following removal, fish assemblage throughout both rivers shifted to a more evenly distributed representation of habitat and reproductive guilds, while restoration of lotic habitat conditions increased, as surface water temperatures decreased and QHEI, IBI, and dissolved oxygen increased. This shift in environmental metrics and increase in overall stream quality increased, particularly in the formerly impounded reaches, indicate diminished habitat homogeneity, and a shift towards natural habitat diversity. This habitat diversification likely led to the restoration of a range of potential niches, thereby increasing the array of guild types inhabiting these rivers, while simultaneously preventing single-guild dominance.     

PREDICTING HABITAT RESPONSE TO FLOW USING GENERALIZED HABITAT MODELS FOR TROUT IN ROCKY MOUNTAIN STREAMS

Dams and water diversions can dramatically alter the hydraulic habitats of stream ecosystems. Predicting how water depth and velocity respond to flow alteration is possible using hydraulic models, such as Physical Habitat Simulation (PHABSIM); however, such models are expensive to implement and typically describe only a short length of stream (10² m). If science is to keep pace with development, then more rapid and cost‐effective models are needed. We developed a generalized habitat model (GHM) for brown and rainbow trout that makes similar predictions to PHABSIM models but offers a demonstrated reduction in survey effort for Colorado Rocky Mountain streams. This model combines the best features of GHMs developed elsewhere, including the options of desktop (no‐survey) or rapid‐survey models. Habitat–flow curves produced by PHABSIM were simplified to just two site‐specific components: (i) Q95h (flow at 95% of maximum habitat) and (ii) Shape. The Shape component describes the habitat–flow curves made dimensionless by dividing flow increments by Q95h and dividing habitat (weighted usable area) increments by maximum habitat. Both components were predicted from desktop variables, including mean annual flow, using linear regression. The rapid‐survey GHM produced better predictions of observed habitat than the desktop GHM (rapid‐survey model explained 82–89% variance for independent validation sites; desktop 68–85%). The predictive success of these GHMs was similar to other published models, but survey effort to achieve that success was substantially reduced. Habitat predicted by the desktop GHM (using geographic information system data) was significantly correlated with the abundance of large brown trout (p < 0.01) but not smaller trout. Copyright © 2013 John Wiley & Sons, Ltd.     

Effects of variation in streamflow and channel structure on smallmouth bass habitat in an alluvial stream

We evaluated the effects of streamflow‐related changes in channel shape and morphology on the quality, quantity, availability and spatial distribution of young‐of‐year and adult smallmouth bass Micropterus dolomieu habitat in an alluvial stream, the Baron Fork of the Illinois River, Oklahoma. We developed Habitat Suitability Criteria (HSC) for young‐of‐year and adult smallmouth bass to assess changes in available smallmouth bass habitat between years, and compare predicted smallmouth bass Weighted Usable Area (WUA) with observed WUA measured the following year. Following flood events between 1999 and 2000, including a record flood, changes in transect cross‐sectional area ranged from 62.5% to 93.5% and channel mesohabitat overlap ranged from 29.5% to 67.0% in study three study reaches. Using Physical HABitat SIMulation (PHABSIM) system analysis, we found that both young‐of‐year and adult smallmouth bass habitat were differentially affected by intra‐ and inter‐annual streamflow fluctuations. Maximum WUA for young‐of‐year and adults occurred at streamflows of 1.8 and 2.3 m³ s^?1, respectively, and WUA declined sharply for both groups at lower streamflows. For most microhabitat variables, habitat availability was similar between years. Habitat suitability criteria developed in 1999 corresponded well with observed fish locations in 2000 for adult smallmouth bass but not for young‐of‐year fish. Our findings suggest that annual variation in habitat availability affects the predictive ability of habitat models for young‐of‐year smallmouth bass more than for adult smallmouth bass. Furthermore, our results showed that despite the dynamic nature of the gravel‐dominated, alluvial Baron Fork, HSC for smallmouth bass were consistent and transferable between years. Published in 2008 by John Wiley & Sons, Ltd.