A LIDAR‐DERIVED EVALUATION OF WATERSHED‐SCALE LARGE WOODY DEBRIS SOURCES AND RECRUITMENT MECHANISMS: COASTAL MAINE,USA

In‐channel large woody debris (LWD) promotes quality aquatic habitat through sediment sorting, pool scouring and in‐stream nutrient retention and transport. LWD recruitment occurs by numerous ecological and geomorphic mechanisms including channel migration, mass wasting and natural tree fall, yet LWD sourcing on the watershed scale remains poorly constrained. We developed a rapid and spatially extensive method for using light detection and ranging data to do the following: (i) estimate tree height and recruitable tree abundance throughout a watershed; (ii) determine the likelihood for the stream to recruit channel‐spanning trees at reach scales and assess whether mass wasting or channel migration is a dominant recruitment mechanism; and (iii) understand the contemporary and future distribution of LWD at a watershed scale. We utilized this method on the 78‐km‐long Narraguagus River in coastal Maine and found that potential channel‐spanning LWD composes approximately 6% of the valley area over the course of the river and is concentrated in spatially discrete reaches along the stream, with 5 km of the river valley accounting for 50% of the total potential LWD found in the system. We also determined that 83% of all potential LWD is located on valley sides, as opposed to 17% on floodplain and terrace surfaces. Approximately 3% of channel‐spanning vegetation along the river is located within one channel width of the stream. By examining topographic and morphologic variables (valley width, channel sinuosity, valley‐side slope) over the length of the stream, we evaluated the dominant recruitment processes along the river and often found a spatial disconnect between the location of potential channel‐spanning LWD and recruitment mechanisms, which likely explains the low levels of LWD currently found in the system. This rapid method for identification of LWD sources is extendable to other basins and may prove valuable in locating future restoration projects aimed at increasing habitat quality through wood additions. Copyright © 2011 John Wiley & Sons, Ltd.     

SPATIAL PATTERN,DENSITY AND CHARACTERISTICS OF LARGE WOOD IN CONNECTICUT STREAMS: IMPLICATIONS FOR STREAM RESTORATION PRIORITIES IN SOUTHERN NEW ENGLAND

Many streams have been modified so extensively that river managers do not have clear reference conditions to frame targets for stream restoration. Large woody debris (LWD) has long been recognized as an important influence on both geomorphic and ecologic processes in stream channels; however, there have been few studies of LWD dynamics in New England. Although this region is heavily forested today, the forest is predominantly young (70–90 years old) regrowth following a historical episode of severe deforestation. This study presents the results of an extensive census of LWD and associated stream characteristics in over 16 river kilometres of northeastern Connecticut streams and represents the first reported inventory of wood loading and sorting in Southern New England. Results of this study indicate that wood loading and jam frequencies in the study region are low: 2.5–17.8 and 0.5–5.51 per 100 m, respectively. Orientation of LWD is predominantly parallel to flow, an indication that these streams are not retaining organic matter or sediment, which has important geomorphic and ecologic implications. Results imply that stream recruitment of LWD is still lagging from the massive forest conversions of the 18th and 19th centuries. Given the low wood loadings observed in the study reaches, manual wood addition and continued forest regeneration would likely improve both habitat diversity and organic matter and fine sediment retention in these systems. Copyright © 2011 John Wiley & Sons, Ltd.     

Effects of large woody debris on surface structure and aquatic habitat in forested streams,southern interior British Columbia,Canada

It is well known that large woody debris (LWD) plays an important functional role in aquatic organisms' life. However, the influence of LWD on channel morphology and aquatic environments at watershed levels is still unclear. The relationships between wood and surface structure and aquatic habitat in 35 first through fifth order streams of southern interior British Columbia were investigated. Study streams in the channel networks of the study watersheds were classified into four size categories based on stream order and bankfull width: Stream size I: bankfull width was less than 3 m, Stream size II: 3–5 m, Stream size III: 5–7 m, Stream size IV: larger than 7 m. We found the number of functional pieces increased with stream size and wood surface area in stream sizes I, II and III (24, 28 and 25 m²/100 m², respectively) was significantly higher than that in stream size IV (12 m²/100 m²). The contribution of wood pieces to pool formation was 75% and 85% in stream sizes II and III, respectively, which was significantly higher than those in stream size I (50%) and size IV (25%). Between 21% and 25% of wood pieces were associated with storing sediment, and between 20% and 29% of pieces were involved in channel bank stability in all study streams. Due to long‐term interactions, LWD in the intermediate sized streams (Size II and III) exhibited much effect on channel surface structure and aquatic habitats in the studied watersheds. Copyright © 2008 John Wiley & Sons, Ltd.     

Quantity,controls and functions of large woody debris in Midwestern USA streams

Large woody debris (LWD) can increase stream habitat heterogeneity by providing structure, altering flow patterns, enhancing sediment deposition, forming pools and retaining organic matter. In North America, the role of LWD has been studied extensively in streams of mature forests (e.g. Pacific Northwest), but few studies have assessed LWD in streams of younger forests (e.g. Midwestern USA). Our objectives were to: (1) quantify the volume and abundance of LWD in a set of Midwestern streams; (2) evaluate possible factors influencing LWD quantity; (3) identify the functional roles of LWD; and (4) compare LWD levels in the upper Midwest to those elsewhere in North America. In 2002 and 2003, we measured LWD and geomorphological variables in 15 low‐gradient streams draining previously logged watersheds in the Upper Peninsula of Michigan. Mean (±SE) LWD volume (0.77 ± 0.12 m³ 100 m⁻²) and abundance (33 ± 3 pieces 100 m⁻¹) were 71% and 10% lesser, respectively, than in streams of similar gradient elsewhere in North America. Channel shape (width:depth ratio) explained 30% of the variation in LWD volume (multiple stepwise regression, P = 0.015) while LWD length and length:channel width combined, explained 72% of the variation in LWD density (multiple stepwise regression, P < 0.0001). About 50% of the LWD either stored sediment or stabilized banks and 14% of the LWD formed pools, although pool density was not significantly related to LWD volume or density. LWD levels, overall, were low in upper Midwestern streams, but the relative importance of that LWD to ecosystem function may be magnified in these wood‐poor systems. Copyright © 2006 John Wiley & Sons, Ltd.     

Flume study of the hydraulic effects of large woody debris in lowland rivers

As a part of a study investigating the hydraulic effects of large woody debris (LWD) in lowland rivers, a series of small-scale experiments were conducted in a rectangular glass-walled recirculating flume. These experiments were undertaken to determine the order of magnitude of the increase in flood levels caused by LWD at different positions within a channel cross-section. Position variables that were considered in these experiments were height above bed, angle to flow direction, and separation distance in the direction of flow. This study was undertaken to quantify the hydraulic benefits (primarily reduced flood levels) gained by the removal of LWD from lowland rivers, which is a common practice in several countries. From an integrated river management perspective it is necessary to weigh any hydraulic benefits of LWD removal up against the environmental costs of loss of faunal habitat, and possible geomorphic instability. The results of these experiments indicate that the levels of LWD commonly occurring in the lowland rivers of southeastern Australia seldom cause any significant effect on flood levels. However, where LWD occur at channel constrictions, or where unusually high densities of LWD are present, the effect on flood levels will be significant.     

Post‐flooding distribution and characteristics of large woody debris piles along the semi‐arid Sabie River,South Africa

The formation of large woody debris (LWD) piles has a profound impact on channel patterns and riparian succession in temperate rivers. The opportunity to study LWD along the Sabie River, a river in the semi‐arid region of Kruger National Park, South Africa, arose in February 2000 after a significant flood (c. 100‐year return interval) removed a large proportion of the fully mature riparian forest and other plant communities. Much of the uprooted vegetation was deposited as LWD piles (woody vegetation accumulations deposited on the ground > 0.1 m³) throughout the riparian and upland zones. In this article we describe the spatial distribution patterns of LWD as related to geomorphic channel type and flood frequency zone, and assess pile composition characteristics six months after the flood. Within the areas surveyed there were 68 LWD piles per hectare, the median size of LWD piles was 4.6 m³ but pile sizes (by volume) varied widely. Pool/rapid geomorphic channel types had the highest density of LWD piles (79 ha^?1) and the largest piles (by volume) were in the bedrock anastomosing channels (mean = 124 m³). Piles were larger in the seasonal and ephemeral flood frequency zones (mean = 54 m³ and 55 m³) than piles in the active zone (c. 2 m³). The patterns of distribution and volume of LWD will affect the subsequent development of vegetation communities as debris piles form a mosaic of patches of surviving organisms and propagules that can strongly influence the initial trajectory of succession. The amount, distribution, and subsequent decomposition of LWD are different from that reported for temperate rivers, suggesting that the role of LWD may be different on non‐floodplain rivers such as the Sabie in semi‐arid South Africa. Copyright © 2004 John Wiley & Sons, Ltd.     

Effects of artificial woody structures on Atlantic salmon habitat and populations in a Nova Scotia stream

A 1‐km reach of Brierly Brook, Nova Scotia, was studied from 1995 to 2004 to determine if the addition of artificial structures mimicking large woody debris could enhance Atlantic salmon populations. In 1995, digger logs (which mimic fallen trees) and deflectors (which narrow the channel) were constructed in a 250‐m section of the brook devoid of woody debris (Old Restored Site). In 2003, 5 more digger logs and defectors were built in a previously unrestored section of the stream (New Restored Site). A third control site was left unchanged. Physical changes caused by the structures were monitored at the New Restored Site. Densities of juvenile and spawning Atlantic salmon were also monitored. At all sites, woody debris structures in the brook were important and effective in creating complex salmonid habitat. The structures narrowed the channel, scoured pools and undercut banks. They created habitat that parr used for summer and winter refuge and adult spawners used for cover and resting during upstream migration and spawning. The structures caused gravels to accumulate that spawning adults used to build redds and fry used for shelter. The reaches with structures had higher spawning densities than reaches without them; spawning increased in the New Restored Site relative to the control site. The absence of woody debris may be a bottleneck for salmonid populations in streams of the Atlantic Northeast. For streams with a small or immature riparian zone and little woody debris in the channel, woody structures may be an effective tool for restoring salmonid populations. Copyright © 2008 John Wiley & Sons, Ltd.     

The Use of Stream Power as an Indicator of Channel Sensitivity to Erosion and Deposition Processes

Stream power is a measure of the main driving forces acting in a channel and determines a river's capacity to transport sediment and perform geomorphic work. Recent digital elevation models allow the calculation of channel gradient and consequently stream power at unprecedented spatial resolution, opening promising and novel opportunities to investigate river geomorphic processes and forms. The present paper investigates the suitability of map‐derived information on total and specific stream power (SSP) to identify dominant processes within the channel (i.e. erosion, transport or deposition). SSP has been already used to identify a threshold for channel stability. This paper tests this knowledge and investigates whether or not attributes of stream power profiles are statistically correlated with distinctive field morphological forms. Two gravel bed single‐thread English rivers are used as case studies, the Lune and the Wye. Available deposition and erosion features surveyed in the field from 124 different locations are used to classify channel reaches as erosion, transport or deposition dominated. Meaningful patterns emerge between the stream power attributes and the field‐based channel classification. An SSP threshold, which erosion is triggered, compares favourably with the ones in the literature. Information about upstream stream power profiles helps to determine the dominant processes. The joint configuration of local and upstream stream power information uniquely classifies reaches into four classes of different sensitivity to erosion and deposition. Copyright © 2013 John Wiley & Sons, Ltd.     

Transient organic jams in Puerto Rican mountain streams after hurricanes

Extreme storms in forested environments commonly increase inputs of coarse particulate organic matter (CPOM) and large wood (LW) to streams. Protruding boulders and bedforms, mid‐channel bars, and standing trees can trap CPOM and LW. These organic accumulations can become large enough to span the bankfull channel width, or the accumulations can be predominantly along the channel margins. We refer to both types of accumulations as transient organic jams (TOJs). TOJs can create diverse geomorphic and ecological effects in channel and floodplain environments. We use data collected from mountain streams of the Luquillo Mountains of north‐eastern Puerto Rico following September 2017 Hurricanes Irma and Maria. We examine the location, characteristics, and geomorphic effects of TOJs in channel segments representing diverse drainage areas and channel gradients. We ask three questions: (a) Does the downstream spacing of TOJs correlate with variables such as drainage area or channel gradient? (b) What variables best predict the volume of organic matter within individual TOJs or within a channel segment? And (c) is there a threshold within a river network that separates channel segments with channel‐spanning versus marginal TOJs? Datasets include multiple TOJs along each of 12 stream segments and presence/absence of channel‐spanning TOJs along an additional six streams. Data analysis with multiple linear regressions indicates that downstream spacing, average volume, and total volume per channel length of TOJs correlate significantly with bankfull channel width. Using the akaike information criterion with correction (AICc) model selection method, Strahler stream order has the most influence on the probability of TOJs being marginal or spanning.     

A new methodology for the assessment of large woody debris accumulations on highly modified rivers (example of two French Piedmont rivers)

Methodologies that have been developed to quantify large woody debris (LWD) have been largely tested and adapted for mountain streams of the Pacific Northwest, characterised by a very high density of LWD, composed of large pieces of wood. In French rivers, LWD studies have focused on larger systems presenting low density and discrete distributions of LWD accumulations, where existing methods could not readily be used. We thus propose an easy-to-use method to quantify LWD within such systems. After defining three representative types of LWD, the volume is obtained by representing each LWD accumulation by a simple geometric form in order to measure its height, width and length. A model is then built for the different accumulation types to estimate wood mass from the measured volume. Since the measured volume is a combination of air and wood, we quantified the proportion of air, which is, respectively, equal to 18, 90 and 93% for trunks, wood jams and shrubs. To understand variability in wood mass, we evaluated the influence of different factors on wood density (defined as the ratio between mass and volume). The main factor was found to be the water absorption capacity of the wood, whereas a lesser factor was the degree of wood decay. Most wood pieces were found to increase their mass by an average of 100% and more after only 24 h in contact with water. Moreover, the observed levels of water loss and water absorption during the first 24 h of removal or exposure to water imply major short-term variations in wood mass, which may have significant consequences for wood transport during flooding. © 1998 John Wiley & Sons, Ltd.     

Geomorphic applications of stream‐gage information

In the United States, several thousand stream gages provide what typically is the only source of continuous, long‐term streamflow and channel‐geometry information for the locations being monitored. In this paper, the geomorphic content of stream‐gage information, previous and potential applications of stream‐gage information in fluvial geomorphic research and various possible limitations are described. Documented applications include studies of hydraulic geometry, channel bankfull characteristics, sediment transport and channel geomorphic response to various types of disturbance. Potential applications include studies to determine the geomorphic effectiveness of large floods and in‐stream habitat change in response to disturbance. For certain applications, various spatial, temporal and data limitations may render the stream‐gage information of limited use; however, such information often is of considerable value to enable or enhance geomorphic investigations. Published in 2008 by John Wiley & Sons, Ltd.     

The development of hydraulic and geomorphic complexity in recently formed streams in Glacier Bay National Park,Alaska

Geomorphic and hydraulic complexity within five streams representing 200 years of stream development were examined in Glacier Bay National Park, Alaska. Channel geomorphic units (CGUs) were mapped using a hierarchical approach, which defined stream habitat according to morphological and hydraulic characteristics. Detailed hydraulic assessment within the geomorphic units allowed differences in hydraulic characteristics across the 200‐year chronosequence to be documented. Channel geomorphology and hydrology changed as stream age increased. Younger streams were dominated by fast flowing geomorphic units such as rapids and riffles with little hydraulic or landscape diversity. As stream age increased, slower flowing habitat units such as glides and pools became more dominant, resulting in increased geomorphic, hydraulic and landscape diversity. These results suggest that geomorphic and hydraulic complexity develop over time, creating habitat features likely to be favoured by instream biota, enhancing biodiversity and abundance. Copyright © 2009 John Wiley & Sons, Ltd.     

LARGE WOODY DEBRIS IN URBAN STREAM CHANNELS: REDEFINING THE PROBLEM

Large woody debris (LWD) is an important ecological element in rivers and streams. Despite its importance, LWD is often removed from urban stream channels for flood control or road maintenance purposes, an approach with high economic and ecological costs and one that is largely unsuccessful. We propose an approach to conserve LWD in channels by modifying infrastructure (culverts and bridges) to allow LWD passage, maintaining aquatic habitat and reducing flooding and road maintenance costs. In Soquel Creek (California, USA), which has a history of LWD‐related flooding, we compared long‐term LWD management costs of historical, current and a LWD‐passing approach whereby infrastructure is enlarged to accommodate LWD passage downstream. We estimated costs of infrastructure replacement, programmatic flood control (LWD removal), LWD‐related flood damage and lost aquatic habitat. The amount of lost aquatic habitat was determined by comparing LWD loading (pieces m^?1) in Soquel Creek (0.007 pieces m^?1) to nearby unmanaged streams (0.054 to 0.106 pieces m^?1). Estimated costs of infrastructure able to pass LWD were nearly double that of historical costs but comparable to current costs. The LWD‐passing approach was comparable to removal approaches in the short term (1 to 50 years) but much less in the long term (51 to 100 years), as expenditures in infrastructure replacement to accommodate LWD yielded reductions in flooding costs and habitat loss. Given the urgency to maintain and restore aquatic habitat, the proposed approach may be broadly applicable. Copyright © 2011 John Wiley & Sons, Ltd.     

Detecting Fluvial Wood in Forested Watersheds using LiDAR Data: A Methodological Assessment

It has long been known that large wood in rivers increases channel complexity and is a primary driver of geomorphic change in forested mountain streams in the Pacific Northwest. Studies analyzing the presence and distribution of fluvial wood are often limited in their spatial extents to the site or reach scales because of the intensive fieldwork required for comprehensive wood surveys. Remote sensing techniques are beginning to allow researchers to assess fluvial wood dynamics and distributions on a basin or regional scale. We used 2009 high‐resolution light detection and ranging (LiDAR) point cloud data to detect and quantify wood within five forested watersheds in the Oregon Coast Range. We filtered the LiDAR data to remove the forest canopy over the stream channel and visually inventoried fluvial wood based on its distinct shape within the channels. We derived several wood and stream morphometric variables to test theories relating to wood abundance and positioning in the lower reaches of streams. We were able to detect fluvial wood with confidence; however, validation of results with ground‐truth data was difficult in the study due to the dynamic and mobile nature of wood through time. We mapped a total of 163 single logs and 55 logjams within the five study watersheds. We did not find statistically significant differences between individual pieces and jam positioning in relation to slope; however, the surveyed wood was often found in areas of lower stream power. This research shows that it is possible to use height‐filtered LiDAR to detect in‐stream wood in densely forested watersheds and has the potential to be employed in future wood studies across broad spatial scales. Copyright © 2015 John Wiley & Sons, Ltd.     

Toledo Bend reservoir and geomorphic response in the lower Sabine River

Downstream geomorphic responses of stream channels to dams are complex, variable, and difficult to predict, apparently because the effects of local geological, hydrological, and operational details confound and complicate efforts to apply models and generalizations to individual streams. This sort of complex geomorphic response characterizes the Sabine River, along the Texas and Louisiana border, downstream of the Toledo Bend dam and reservoir. Toledo Bend controls the flow of water and essentially prevents the flux of sediment from three‐quarters of the drainage basin to the lower Sabine River. Although the channel is scoured immediately downstream of the dam, further downstream there is little evidence of major changes in sediment transport or deposition, sand supply, or channel morphology attributable to the impoundment. Channels are actively shifting, banks are eroding, and sandbars are migrating, but not in any discernibly different way than before the dam was constructed. The Sabine River continues to transport sand downstream, and alluvial floodplains continue to accrete. The relatively small geomorphic response can be attributed to several factors. While dam releases are unnaturally flashy and abrupt on a day‐to‐day basis, the long‐term pattern of releases combined with some downstream smoothing creates a flow regime in the lower basin which mimics the pre‐dam regime, at least at monthly and annual time scales. Sediment production within the lower Sabine basin is sufficient to satisfy the river's sediment transport capacity and maintain pre‐dam alluvial sedimentation regimes. Toledo Bend reservoir has a capacity: annual inflow ratio of 1.2 and impounds 74% of the Sabine drainage basin, yet there has been minimal geomorphic response in the lower river, which may seem counterintuitive. However, the complex linked geomorphic processes of discharge, sediment transport and loads, tributary inputs, and channel erosion include interactions which might increase as well as decrease sediment loads. Furthermore, if a stream is transport‐limited before impoundment, the reduced sediment supply after damming may have limited impact. Copyright © 2003 John Wiley & Sons, Ltd.     

Invertebrate communities in Compensation Creek,a man‐made stream in boreal Newfoundland: The influence of large woody debris

Benthic invertebrate communities were examined in Compensation Creek, a man‐made stream in south‐central Newfoundland, Canada. Samples taken in September 2006 and September 2007 from large woody debris (LWD) were compared with samples from benthic environments to determine whether LWD supported a more diverse and abundant invertebrate community. Benthic habitats in a nearby natural stream were also sampled. Taxa composition was similar between the man‐made and the natural stream, highlighting successful colonization for the majority of taxa. Within Compensation Creek, taxa richness was higher in benthic habitats than on LWD, likely influenced by the successional age of the stream and surrounding habitat. The more complex benthic substrate provided refugia and allowed for the accumulation of fine detritus as a food source. Scrapers were almost completely absent from LWD and collector‐gatherer abundance was greater in the benthos. Collector‐filterer abundance was more than six times greater near the pond outflow than farther downstream when discharge was high, but abundances were almost equal when discharge was reduced. Riparian vegetation has not fully established around the man‐made stream, whereas it is overhanging and extensive at the natural stream, leading to more leaf‐litter input for shredders. As the morphology of Compensation Creek changes, the invertebrate community will continue to develop and likely increase utilization of accumulated detritus at LWD. Copyright © 2009 John Wiley & Sons, Ltd.     

Lowland stream restoration by sand addition: Impact,recovery, and beneficial effects on benthic invertebrates

Up to now, most lowland stream restoration projects were unsuccessful in terms of ecological recovery. Aiming to improve the success of stream restoration projects, a novel approach to restore sandy‐bottom lowland streams degraded by channel incision was launched, consisting of the addition of sand to the stream channel in combination with the introduction of coarse woody debris. Yet it remained unknown whether this novel measure of sand addition is actually effective in terms of biodiversity improvements. The aim of the present study was therefore to evaluate if sand addition can improve hydromorphological stream complexity on the short term leading to an increase in macroinvertebrate biodiversity. To this end, particle transport, water depth, current velocity, dissolved oxygen dynamics, and sediment composition were measured. The response of the macroinvertebrate community composition was determined at different stages during the disturbance and short‐term recovery process. Immediately downstream the sand addition site, transport and sedimentation of the sand were initially intense, until an equilibrium was reached and the physical conditions stabilized. The stream section matured fast as habitat formation took place within a short term. Macroinvertebrate diversity decreased initially but recovered rapidly following stabilization. Moreover, an increase in rheophilic taxa was observed in the newly formed habitats. Thus, although sand addition initially disturbed the stream, a relatively fast physical and biological recovery occurred, leading to improved instream conditions for a diverse macroinvertebrate community, including rheophilic taxa. Therefore, we concluded that sand addition is a promising restoration measure for incised lowland streams.     

Spatial distribution of large wood jams in streams related to stream‐valley geomorphology and forest age in Northern Michigan

Geomorphology at the scale of stream valleys influences smaller scale processes that give rise to spatially distributed patches, including large wood jams (LWJ) in streams. Understanding the spatial distribution of LWJ along streams with reference to large‐scale geomorphology is valuable for understanding stream and riparian interactions, and may be critical for effective stream management and restoration. We surveyed the locations of LWJ along 18 stream segments within study areas defined by stream‐valley geomorphology. The objective of this study was to test the prediction that LWJ in this system will be aggregated in areas defined by stream‐valley geomorphology, but be randomly distributed at smaller scales. The spatial distribution of LWJ was analysed by a one‐dimensional K‐function analysis capable of detecting aggregated, random and segregated patterns at different scales. The prediction that LWJ aggregate in areas defined by stream‐valley geomorphology was supported: LWJ aggregated at scales up to several kilometres in three streams. LWJ also was segregated at smaller scales in two of these streams; this was detectable when several stream valley segments were considered together. The prediction that LWJ would be randomly distributed at smaller scales was supported at most smaller scales for most streams. In fact, 40% of individual stream valley segments contained LWJ that were randomly distributed at all scales. Twenty‐seven per cent of individual stream valley segments showed segregated LWJ distributions. Large‐scale aggregation of LWJ evidences the need to select reference reaches that encompass several geomorphic patches at the scale of the stream valley. Segregated patterns of LWJ distributions evidence opportunities to better understand the relationships between hydraulic systems, material transport dynamics and riparian forests. Copyright © 2009 John Wiley & Sons, Ltd.     

Macroinvertebrate community structure and function associated with large wood in low gradient streams

Large woody debris (wood) plays a number of important roles in forested stream ecosystems. Wood in streams provides habitat and flow refugia for fish and invertebrates, and is a site of biofilm production that serves as food for grazing organisms. Logs added to streams are rapidly colonized by invertebrates, and this habitat alteration is accompanied by changes in community composition and functional processes. A multiple habitat, qualitative sampling approach was employed to evaluate macroinvertebrate communities associated with wood habitats in 71 stream reaches in central Michigan and southeastern Minnesota, two agricultural regions in the midwestern United States. Macroinvertebrate taxa were classified with respect to behaviour (e.g. sprawler, clinger, swimmer), as well as trophic/feeding characteristics. These traits were used to examine community structure as a function of wood abundance and distribution. Although wood is not abundant in these streams and logs are generally small in size, wood is a very important habitat in both Michigan and Minnesota: 86% and 95% of the total taxa encountered at Michigan and Minnesota study sites, respectively, were found in wood habitats. Differences in regional patterns in the distribution of taxa across habitats were observed between Michigan and Minnesota. These are believed to result from differences in the number of habitat types available, and the dominant substrate composition. Local invertebrate diversity increased in Michigan, but not Minnesota, with the presence of wood habitats in streams. The presence of wood at a site increased the average taxa richness by 15 and 10 taxa in Michigan and Minnesota, respectively. Macroinvertebrate behavioural attributes and functional traits associated with wood habitats suggest that community traits may vary due to both local difference in flow and the location of wood in the channel. Copyright © 2003 John Wiley & Sons, Ltd.     

The role of riparian vegetation and woody debris in the development of macroinvertebrate assemblages in streams

Riparian vegetation development and macroinvertebrate assemblages were studied in 16 streams formed between 35 and 230 years ago, following glacial recession in Glacier Bay National Park, southeast Alaska. Riparian vegetation established most rapidly in streams where flow variation in downstream reaches was buffered by a lake. Riparian vegetation development was positively correlated with lower bank stability, but was independent of stream age. Roots and branches of riparian vegetation trailing into streams (trailing riparian habitat—TRH) were shown to be an important habitat for a number of macroinvertebrate taxa. In young and unstable streams, TRH was colonized mainly by Plecoptera whereas in more stable lake‐influenced streams Simuliidae dominated. Significant coarse woody debris (CWD) accumulations were not observed until after approximately 130 years of stream development had occurred when certain channel features, such as gravel bars, were stabilized by dead wood. Where dead wood was present, opportunistic wood taxa were abundant, even in the younger streams. However, a xylophagous species, Polypedilum fallax, was not recorded until streams were over 100 years old. Two‐way indicator species analysis (TWINSPAN) using presence/absence of macroinvertebrate taxa on TRH, initially divided streams into lake and non‐lake systems, but subsequent divisions were consistent with differences in stream age. TWINSPAN of macroinvertebrate assemblages on dead wood again highlighted differences in stream age. Canonical correspondence analysis indicated that bed stability and stream age were the most important environmental variables influencing macroinvertebrate distribution on TRH. Trailing riparian habitat was most abundant in moderately unstable streams where it facilitates invertebrate colonization. CWD contributes markedly to channel stabilization, provides habitat for invertebrate xylophages, and confers additional habitat complexity. Maximum levels of CWD are predicted to occur in non‐lake streams after approximately 300 years, but at least a further 100 years will be required in stable streams below lakes where dead wood entrainment is not enhanced by flooding, channel migration and bank undercutting. A conceptual model summarizing the role of TRH and CWD on stream development in Glacier Bay is presented. Copyright © 2005 John Wiley & Sons, Ltd.