DISCONNECTING THE FLOODPLAIN: EARTHWORKS AND THEIR ECOLOGICAL EFFECT ON A DRYLAND FLOODPLAIN IN THE MURRAY–DARLING BASIN,AUSTRALIA

Globally, dams and water extractions are well‐recognised disruptors of flow regimes in floodplain wetlands, but little is known of the hydrological and ecological impacts of floodplain earthworks constructed for irrigation, flood mitigation and erosion control. We mapped the distribution of earthworks with high‐resolution SPOT (Système Probatoire d'Observation de la Terre) imagery in an internationally recognised Ramsar wetland, the Macquarie Marshes of the Murray–Darling Basin, Australia. There were 339 km levees, 1648 km channels, 54 off‐river storages and 664 tanks (0.5–5 m high), detected within the 4793 km² floodplain study area. Earthworks reduced localised flooding compared with undeveloped sites. The most pronounced disconnection of the original floodplain (73.0%) occurred where earthworks were most concentrated compared with areas with few earthworks (53.2%). We investigated relationships between hydrological connectivity and mortality of the perennial flood‐dependent river red gum Eucalyptus camaldulensis at 55 floodplain sites (225 × 150 m). Over half of the river red gums were dead at 21.8% of the sites. Earthworks blocked surface flows to flood‐dependent vegetation and drowned vegetation in artificially inundated off‐river storages. Mortality was due to impacts of earthworks and potentially exacerbated by effects of river regulation, water extraction and climate. River red gums were healthiest in narrow river corridors where earthworks confined flows and flows could recede freely. Rehabilitation of flood‐dependent ecosystems should focus on reinstating lateral connectivity and protecting environmental flows. Copyright © 2011 John Wiley & Sons, Ltd.     

VEGETATION–LANDFORM ASSEMBLAGES ALONG SELECTED RIVERS IN THE CZECH REPUBLIC,A DECADE AFTER A 500‐YEAR FLOOD EVENT

Rivers with a natural flow regime strongly influence the dynamics of riparian plant communities through hydrological and geomorphological processes. In this study, associations between fluvial landforms and vegetation are investigated on three near‐natural rivers in the Czech Republic a decade after a 500‐year return period flood in July 1997. This extreme disturbance destroyed the anthropogenically modified river channels and created suitable conditions for a range of ecosystems with high diversity and ecological stability. Field surveys were conducted on fluvial landforms (bars, islands, banks, floodplains and terraces) along three ‘renaturalized’ rivers, where no technical modifications had subsequently been made to their channels outside urban areas and the floodplains had been left in a post‐flood state. Vegetation species abundance and 13 environmental variables (topographical, hydrological and soil) were investigated in summer 2007, 10 years after the extreme flood disturbance. The results suggest that the recently created fluvial geomorphic forms are key environmental determinants of riparian vegetation distribution patterns. A range of statistical analyses illustrate that some plant species show predictable patterns of occurrence that correspond with the fluvial forms, supporting a fourfold grouping of herbaceous and woody species and the identification of typical plant communities associated with gravel bars, islands, banks, floodplains and terraces. An investigation of the species richness found on different fluvial landforms showed that the highest number of species occurred on the floodplain and decreased gradually towards the channel bed and towards terraces. Investigation of existing conditions in reaches of rivers with natural dynamics of fluvial processes provides valuable information that can be used as an effective tool for planning restoration strategies and precise management. However, the most important finding of this study is the remarkable establishment of complex river corridor vegetation–landform associations within 10 years of a 500‐year flood that removed the heavily cultivated landscape that had existed before the event. Copyright © 2011 John Wiley & Sons, Ltd.     

SEDIMENT DYNAMICS IN THE RESTORED REACH OF THE KISSIMMEE RIVER BASIN,FLORIDA: A VAST SUBTROPICAL RIPARIAN WETLAND

Historically, the Kissimmee River Basin consisted of a broad nearly annually inundated riparian wetland similar in character to tropical Southern Hemisphere large rivers. The river was channelized in the 1960s and 1970s, draining the wetland. The river is currently being restored with over 10 000 hectares of wetlands being reconnected to 70 river km of naturalized channel. We monitored riparian wetland sediment dynamics between 2007 and 2010 at 87 sites in the restored reach and 14 sites in an unrestored reference reach. Discharge and sediment transport were measured at the downstream end of the restored reach. There were three flooding events during the study, two as annual flood events and a third as a greater than a 5‐year flood event. Restoration has returned periodic flood flow to the riparian wetland and provides a mean sedimentation rate of 11.3 mm per year over the study period in the restored reach compared with 1.7 mm per year in an unrestored channelized reach. Sedimentation from the two annual floods was within the normal range for alluvial Coastal Plain rivers. Sediment deposits consisted of over 20% organics, similar to eastern blackwater rivers. The Kissimmee River is unique in North America for its hybrid alluvial/blackwater nature. Fluvial suspended‐sediment measurements for the three flood events indicate that a majority of the sediment (70%) was sand, which is important for natural levee construction. Of the total suspended sediment load for the three flood events, 3%–16% was organic and important in floodplain deposition. Sediment yield is similar to low‐gradient rivers draining to the Chesapeake Bay and alluvial rivers of the southeastern USA. Continued monitoring should determine whether observed sediment transport and floodplain deposition rates are normal for this river and determine the relationship between historic vegetation community restoration, hydroperiod restoration, and sedimentation. Published in 2011 by John Wiley & Sons, Ltd.     

FLOOD INUNDATION MAPPING FOR INTEGRATED FLOODPLAIN MANAGEMENT: UPPER MISSISSIPPI RIVER SYSTEM

Natural hydrogeomorphic characteristics and hydrologic alterations are important ecological drivers, and hydrology is also a common ecological, flood control and navigation system indicator. Hydrologic characteristics change dramatically from one end of the Upper Mississippi River System to the other, and hydraulic characteristics also differ spatially across the river channels and floodplain in response to dams, levees and diversions. Low flow surface water spatial change in response to navigation and flood control has been well known for many years, but little information was available on the spatial distribution of frequent floods. The flow frequency data presented here were developed to better estimate contemporary floods after historic flooding in 1993. Flood stage estimates are enhanced in GIS to help quantify and map potential floodplain inundation for more than 1000 river miles on the Upper Mississippi and Illinois Rivers. Potential flood inundation is mapped for the 50% to 0.2% annual exceedance probability flood stage (i.e. 2‐ to 500‐year expected recurrence interval flood) and also for alternative floodplain management scenarios within the existing flood protection infrastructure. Our analysis documents: (i) impoundment effects, (ii) a hydrologic gradient within the navigation pools that creates repeating patterns of riverine, backwater and impounded aquatic habitat conditions, (iii) potential floodplain inundation patterns for over 2 million acres and (iv) several integrated floodplain management scenarios. Extreme flood events are more common in recent decades, and they are expected to continue to occur at greater frequency in response to climate change. Floodplain managers can use the results presented here to help optimize land management and flood damage reduction on the Upper Mississippi River System. Published 2012. This article is a U.S. Government work and is in the public domain in the USA.     

IMPORTANCE OF FLOODPLAIN CONNECTIVITY TO FISH POPULATIONS IN THE APALACHICOLA RIVER,FLORIDA

Floodplain habitats provide critical spawning and rearing habitats for many large‐river fishes. The paradigm that floodplains are essential habitats is often a key reason for restoring altered rivers to natural flow regimes. However, few studies have documented spatial and temporal utilization of floodplain habitats by adult fish of sport or commercial management interest or assessed obligatory access to floodplain habitats for species' persistence. In this study, we applied telemetry techniques to examine adult fish movements between floodplain and mainstem habitats, paired with intensive light trap sampling of larval fish in these same habitats, to assess the relationships between riverine flows and fish movement and spawning patterns in restored and unmodified floodplain distributaries of the Apalachicola River, Florida. Our intent is to inform resource managers on the relationships between the timing, magnitude and duration of flow events and fish spawning as part of river management actions. Our results demonstrate spawning by all study species in floodplain and mainstem river habitat types, apparent migratory movements of some species between these habitats, and distinct spawning events for each study species on the basis of fish movement patterns and light trap catches. Additionally, Micropterus spp., Lepomis spp. and, to a lesser degree, Minytrema melanops used floodplain channel habitat that was experimentally reconnected to the mainstem within a few weeks of completing the restoration. This result is of interest to managers assessing restoration activities to reconnect these habitats as part of riverine restoration programmes globally. Copyright © 2012 John Wiley & Sons, Ltd.     

Monitoring of environmental flow outcomes in a large river basin: The Commonwealth Environmental Water Holder's long‐term intervention in the Murray–Darling Basin,Australia

The Murray–Darling Basin in south‐eastern Australia contains over 70,000 km² of wetlands and floodplains, many of which are in poor condition. In response, Australian governments have committed to a major restoration program, the Murray–Darling Basin Plan that includes management of 2,750 Gl of environmental water to protect and restore aquatic ecosystems. The restoration is being undertaken within an adaptive management framework that includes monitoring the outcomes of environmental flows in seven river valleys. This paper provides an overview of the 5‐year monitoring project and some preliminary results. Monitoring design considered the Basin Plan's environmental objectives, conceptual models of ecosystem responses to flow, and an outcomes framework linking flow responses to the environmental objectives. Monitoring indicators includes ecosystem type, vegetation, river metabolism, and fish. Responses are evaluated to identify the contribution of environmental flows to Basin Plan environmental objectives and continual improvements in management. The program is unique in that it seeks to monitor long‐term outcomes of environmental flows at the river basin scale. Despite many challenges, the monitoring has become a key part of the adaptive management of environmental flows in the Murray–Darling Basin.     

Impact of a 100‐year flood on vegetation,benthic invertebrates,riparian fauna and large woody debris standing stock in an alpine floodplain

This paper investigates the impact of a 100‐year flood in May 1999 on community composition and large woody debris standing stock in an alpine floodplain (Isar, Germany). Detailed pre‐flood data sampled from 1993 to 1998 are compared with the situation directly after the flood. In those parts of the Isar floodplain mainly covered with pioneer vegetation prior to flooding, the coverage of unvegetated gravel bars increased by 22% following the flood. However, the flood did not remove larger amounts of older successional vegetation stages (willow thickets, floodplain forest). No significant changes in the benthic invertebrate fauna were recorded. The lowest densities of riparian ground beetles (Carabidae) within the study were recorded one month after the flood. Two months later, the ground beetle densities increased to the highest values ever recorded, indicating the ground beetle's high potential for recolonization. These results highlight the degree of resilience of both the aquatic and the riparian invertebrate fauna. The flood also caused a significant increase in large woody debris standing stock; in one section the number of logs increased tenfold and the volume increased by a factor of 20, leading to the assumption that most woody debris in alluvial flood‐plains is provided by catastrophic events. Copyright © 2004 John Wiley & Sons, Ltd.     

POST‐DAMMING FLOW REGIME DEVELOPMENT IN A LARGE LOWLAND RIVER (VOLGA,RUSSIAN FEDERATION): IMPLICATIONS FOR FLOODPLAIN INUNDATION AND FISHERIES

Periodic flooding plays a key role in the ecology of floodplain rivers. Damming of such rivers can disturb flooding patterns and have a negative impact on commercial fish yield. The Volga River, the largest river in Europe, has a regulated flow regime after completion of a cascade of dams. Here, we study effects of damming on long‐term discharge variability and flood pulse characteristics. In addition, we evaluate the effects of the altered flood pulse on floodplain ecosystem functioning and commercial fish yields. Our results indicate that both flood pulse and fish populations of the Volga–Akhtuba floodplain have varied considerably over the past decades. After damming, annual maximum peak discharges have decreased, minimum discharges increased, but average discharges remained similar to pre‐damming conditions. Moreover, because of bed level incision of over 1.5 m, a higher discharge is needed to reach bankfull level and inundate the floodplains. Despite this significantly altered hydrological regime and subsequent morphological changes, current discharge management still provides significant spring flooding. However, commercial fish catches did decrease after damming, both in the main channel and in the floodplain lakes. All catches were dominated by species with a eurytopic flow preference, although catches from the main channel contained more rheophilic species, and floodplain catches contained more limnophilic and phytophilic species. The strong increase of opportunistic gibel carp (Carassius gibelio) around 1985 was apparent in the main channel and the floodplain lakes. Despite the hydrological changes, the decrease in overall catches, and the upsurge of gibel, we found a strong positive effect of flood magnitude in the previous year on commercial fish yield in the floodplain lakes. This suggests that under the current discharge management there still is an increased fish growth and/or survival during high floods and that functioning of the floodplain is at least partly intact. Copyright © 2011 John Wiley & Sons, Ltd.     

Legacies of flood reduction on a Dryland river

The Bill Williams (Arizona) is a regulated dryland river that is being managed, in part, for biodiversity via flow management. To inform management, we contrasted riparian plant communities between the Bill Williams and an upstream free‐flowing tributary (Santa Maria). Goals of a first study (1996–1997) were to identify environmental controls on herbaceous species richness and compare richness among forest types. Analyses revealed that herbaceous species richness was negatively related to woody stem density, basal area and litter cover and positively related to light levels. Introduced Tamarix spp. was more frequent at the Bill Williams, but all three main forest types (Tamarix, Salix/Populus, Prosopis) had low understory richness, as well as high stem density and low light, on the Bill Williams as compared to the Santa Maria. The few edaphic differences between rivers (higher salinity at Bill Williams) had only weak connections with richness. A second study (2006–2007) focused on floristic richness at larger spatial scales. It revealed that during spring, and for the study cumulatively (spring and fall samplings combined), the riparian zone of the unregulated river had considerably more plant species. Annuals (vs. herbaceous perennials and woody species) showed the largest between‐river difference. Relative richness of exotic (vs. native) species did not differ. We conclude that: (1) The legacy of reduced scouring frequency and extent at the Bill Williams has reduced the open space available for colonization by annuals; and (2) Change in forest biomass structure, more so than change in forest composition, is the major driver of changes in plant species richness along this flow‐altered river. Our study informs dryland river management options by revealing trade‐offs that exist between forest biomass structure and plant species richness. Copyright © 2010 John Wiley & Sons, Ltd.     

RIVERBED DIGITAL ELEVATION MODELS AS A TOOL FOR HOLISTIC RIVER MANAGEMENT: MOTUEKA RIVER,NELSON, NEW ZEALAND

River management in New Zealand's laterally active gravelly rivers has permitted floodplain development and protection of agricultural resources and infrastructure. Management of these dynamic systems has been hailed as a success for the approaches adopted, namely straightening and confining the river using bank protection and managing riverbed levels by gravel extraction. However, this activity also impacts river morphological/habitat diversity and potential gravel resource, by replacing broad riparian corridors with narrower channels and reducing lateral connectivity with the floodplain. This paper quantifies river behaviour in three laterally confined reaches in the upper Motueka River over a 7‐year period, using annual high‐resolution ground surveys to address the nature of morphological change and associated sediment flux in these reaches with a view to informing management of the gravel resource. Surveys between 2004 and 2010 acquired data to construct digital elevation models (DEMs) of the active riverbed in three ~1‐km‐long reaches. Morphological budgeting based on differencing between successive DEM surfaces reveals complex spatial and temporal patterns of erosion and deposition, demonstrating complex reach dynamics. Overall, volumetric changes suggest these narrowed reaches have been net exporters of sediment, associated with continued channel degradation. This has left bar features, traditionally the focus of gravel extraction in the reaches, relatively isolated from all but extreme flows, limiting replenishment of the gravel resource. The paper demonstrates the utility of riverbed DEMs as a potential tool to frame river character and behaviour at the reach scale in gravel‐bed rivers, thereby providing an important contribution to holistic river management in these systems. Copyright © 2012 John Wiley & Sons, Ltd.     

Large dam removal and early spontaneous riparian vegetation recruitment on alluvium in a former reservoir: Lessons learned from the pre‐removal phase of the Sélune River project (France)

The restoration of ecological continuity along the Sélune River (Normandy, France) involves the removal of two tall hydroelectric dams (36 m removed in 2019 and 16 m in 2021), a project without precedent in Europe. During the pre‐removal phase (2014–2018), we performed scientific monitoring of the vegetation that was colonizing alluvium in the former dam reservoir (length: 19 km; surface area: 151 ha). Our study aimed to analyse if spontaneous vegetation could ecologically restore the riparian zone and help maintain fine sediment after dam removal. We used colonization indicators related to vegetation structure, taxonomic richness and diversity, and composition. These indicators were calculated at two spatial scales (local, at a single site, and broad, along the reservoir). The aim was to (a) characterize the spontaneously established species pool; (b) analyse longitudinal patterns in vegetation colonization; and (c) assess temporal changes in the species community. Our results show that diverse plant communities have developed. Slight differences in longitudinal and lateral patterns existed; they were linked with habitat heterogeneity and the reservoir's slow pace of draining. We observed fast spontaneous terrestrialization, which has resulted in cover stabilization, decreased diversity, and the development of herbaceous riverbank communities, with very few invasive species. This finding suggests stabilization potential is high and passive ecological restoration could occur, at least locally. Further analyses focusing on functional traits could help inform future management decisions regarding revegetation on reservoir alluvium.     

HYDROLOGIC CONNECTIVITY OF FLOODPLAINS,NORTHERN MISSOURI—IMPLICATIONS FOR MANAGEMENT AND RESTORATION OF FLOODPLAIN FOREST COMMUNITIES IN DISTURBED LANDSCAPES

Hydrologic connectivity between the channel and floodplain is thought to be a dominant factor determining floodplain processes and characteristics of floodplain forests. We explored the role of hydrologic connectivity in explaining floodplain forest community composition along streams in northern Missouri, USA. Hydrologic analyses at 20 streamgages (207–5827 km² area) document that magnitudes of 2‐year return floods increase systematically with increasing drainage area whereas the average annual number and durations of floodplain‐connecting events decrease. Flow durations above the active‐channel shelf vary little with increasing drainage area, indicating that the active‐channel shelf is in quasi‐equilibrium with prevailing conditions. The downstream decrease in connectivity is associated with downstream increase in channel incision. These relations at streamflow gaging stations are consistent with regional channel disturbance patterns: channel incision increases downstream, whereas upstream reaches have either not incised or adjusted to incision by forming new equilibrium floodplains. These results provide a framework to explain landscape‐scale variations in composition of floodplain forest communities in northern Missouri. Faust ( 2006 ) had tentatively explained increases of flood‐dependent tree species, and decreases of species diversity, with a downstream increase in flood magnitude and duration. Because frequency and duration of floodplain‐connecting events do not increase downstream, we hypothesize instead that increases in relative abundance of flood‐dependent trees at larger drainage area result from increasing size of disturbance patches. Bank‐overtopping floods at larger drainage area create large, open, depositional landforms that promoted the regeneration of shade‐intolerant species. Higher tree species diversity in floodplains with small drainage areas is associated with non‐incised floodplains that are frequently connected to their channels and therefore subject to greater effective hydrologic variability compared with downstream floodplains. Understanding the landscape‐scale geomorphic and hydrologic controls on floodplain connectivity provides a basis for more effective management and restoration of floodplain forest communities. Published 2013. This article is a U.S. Government work and is in the public domain in the USA.     

ASSESSMENT OF CONSEQUENCES OF SEDIMENT DEFICIT ON A GRAVEL RIVER BED DOWNSTREAM OF DAMS IN RESTORATION PERSPECTIVES: APPLICATION OF A MULTICRITERIA,HIERARCHICAL AND SPATIALLY EXPLICIT DIAGNOSIS

As regards river restoration, it is fundamental to better link human pressures and environmental responses and to take into consideration not only target species or habitat but diverse ecological elements. This permits to assess sustainable restoration plan, especially concerning sediment augmentation below dams. The use of a hierarchical multicriteria approach on the Ain River permits us to assess a diagnosis of sediment deficit impact integrating several morphological (channel shifting, river bed degradation and river bed coarsening) and ecological components (Riparian and floodplain lake and fish communities). Our diagnosis also integrates a temporal and spatial approach better to link human pressures and environmental responses and to identify the dam effects amongst other drivers (e.g. grazing decline and channel regulation). The results confirm causality links between sediment deficit and slight channel bed degradation (0.01 m.year^?1) or channel bed paving and thus highlight the impact of the dam on the drying of the riparian forest and on former channel community. However, the relationship between incision and reduction in active channel lateral mobility is more difficult to establish. The role of sediment deficit in the current variability of the riparian regeneration capacity and, thereby, landscape diversity along the lower valley remains unclear. This study also confirms the relevance of using different ecological indicators, notably because all components present different adjustment time scales, whereas some of them are more sensitive to other impacts. Copyright © 2013 John Wiley & Sons, Ltd.     

FISH RECRUITMENT IS INFLUENCED BY RIVER FLOWS AND FLOODPLAIN INUNDATION AT APALACHICOLA RIVER,FLORIDA

High human demand for limited water resources often results in water allocation trade‐offs between human needs and natural flow regimes. Therefore, knowledge of ecosystem function in response to varying streamflow conditions is necessary for informing water allocation decisions. Our objective was to evaluate relationships between river flow and fish recruitment and growth patterns at the Apalachicola River, Florida, a regulated river, during 2003–2010. To test relationships of fish recruitment and growth as responses to river discharge, we used linear regression of (i) empirical catch in fall, (ii) back‐calculated catch, via cohort‐specific catch curves, and (iii) mean total length in fall of age 0 largemouth bass Micropterus salmoides, redear sunfish Lepomis microlophus and spotted sucker Minytrema melanops against spring–summer discharge measures in Apalachicola River. Empirical catch rates in fall for all three species showed positive and significant relationships to river discharge that sustained floodplain inundation during spring–summer. Back‐calculated catch at age 0 for the same species showed positive relationships to discharge measures, but possibly because of low sample sizes (n = 4–6), these linear regressions were not statistically significant. Mean total length for age 0 largemouth bass in fall showed a positive and significant relationship to spring–summer discharge; however, size in fall for age 0 redear sunfish and spotted sucker showed no relation to spring–summer discharge. Our results showed clear linkages among river discharge, floodplain inundation and fish recruitment, and they have implications for water management and allocation in the Apalachicola River basin. Managed flow regimes that reduce the frequency and duration of floodplain inundation during spring–summer will likely reduce stream fish recruitment. Copyright © 2012 John Wiley & Sons, Ltd.     

Patterns in groundwater nitrogen concentration in the riparian zone of a large semi‐arid river (River Murray,Australia)

Lateral exchanges of surface water between river channels and their floodplains are important for vegetation health and aquatic food‐web productivity in semi‐arid ecosystems. However, the significance of the lateral connectivity via sub‐surface pathways in these systems is not as well understood. Patterns in nitrogen concentration in groundwater and in the unsaturated zone were used to infer the sub‐surface biogeochemistry of N in the riparian zone of a large semi‐arid floodplain (Hattah‐Kulkyne National Park) of the River Murray, Australia. The riparian zone plays a special role in this system as it is an area of transition between fresh surface waters and saline floodplain groundwater. The river was losing water to the floodplain during baseflow conditions but gradients were temporarily reversed following floods. In general, the redox conditions were sub‐oxic to anoxic in riparian groundwater and the main forms of N present were NH and dissolved organic N. There was a gradient in NH concentration from the river to the floodplain, suggesting that the main source of NH was from the decomposition of organic matter in fluvial sediments. Elevated concentrations of NO were occasionally found in shallow groundwater away from the river following floods but tended not to persist. The source of the NO appeared to be unsaturated‐zone NO displaced to the water table during floods. Assuming that denitrification was the main attenuation process, this displacement of unsaturated zone NO to anoxic groundwater could be a significant N removal process from the ecosystem (estimated at 18 kg N ha^?1 for the largest flood during the study). Understanding the impact of river regulation on floodplain nutrient cycles in River Murray floodplains will be challenging because the changes in floodplain hydrology are complex and coincide with salinization of soils and groundwater. Copyright © 2005 John Wiley & Sons, Ltd.     

INTRA‐ANNUAL VARIATION IN RIVER–RESERVOIR INTERFACE FISH ASSEMBLAGES: IMPLICATIONS FOR FISH CONSERVATION AND MANAGEMENT IN REGULATED RIVERS

While much is known about the fish assemblages, habitats, and ecology of rivers and reservoirs, there has been limited study of the fish assemblages in transitional habitats between these lotic and lentic habitats. Data about these river–reservoir interface (RRI) fish assemblages are needed to guide integrated management efforts of river–reservoir ecosystems. The aim of these efforts is to recommend flows for natural river function, conserve native riverine fish assemblages, and maintain reservoir sport fisheries. We used a multigear approach to assess the fish assemblages of four RRIs in the Colorado River Basin, Texas. In addition to characterizing RRI fish assemblages using species richness and evenness metrics, and habitat‐use guilds, we used a multivariate approach to evaluate intra‐annual shifts in species composition and abundance. All RRIs had high species richness and evenness values and included both macrohabitat generalist and fluvial species. RRIs also contained high proportions of the fish species available within each river–reservoir ecosystem, ranging from 55% to 80%. Observed intra‐annual shifts in RRI fish assemblages resulted from changes in abundance of dominant species rather than changes in species composition, with abundance of most species increasing from early spring to summer. Fish species responsible for intra‐annual shifts included mostly floodplain and migratory species, suggesting that species both used littoral habitats within RRIs and migrated through RRIs to river and reservoir habitats. The diversity of fishes found within RRIs highlights the importance of including these areas in future conservation and management efforts of river–reservoir ecosystems. Copyright © 2013 John Wiley & Sons, Ltd.     

GEOMORPHOLOGICAL CONTROLS ON VEGETATION RESPONSES TO FLOW ALTERATIONS IN A MEDITERRANEAN STREAM (CENTRAL‐WESTERN SPAIN)

The expected recovery of the natural conditions of large regulated rivers over the distance downstream from a dam is limited by relative tributary size according to the Serial Discontinuity Concept; however, geomorphology may also influence the recovery process. We examined the woody vegetation of the riparian zone in seven river segments distributed along the regulated reach of the Tiétar River in central‐western Spain, which flows through two distinct geomorphic templates. Whereas the annual runoff has decreased by 30% on average along the entire studied reach following the construction of the Rosarito Dam and the initiation of field irrigation in the region, the magnitude and frequency of the peak flows decreased by 30% immediately downstream from the dam but recovered the natural values with the distance downstream. We evaluated the recovery patterns toward the natural riparian conditions by comparing woody species composition, diversity and distribution of vegetation patches established prior to and after dam completion. Our results did not indicate a recovery gradient of any of the analysed vegetation attributes downstream from the dam. Instead, we found that the difference in the slope of the stream channel and banks, the width of the valley and the size of substratum particles among the surveyed patches were factors that significantly mediated dam and tributary effects on vegetation and influenced the degree of vegetation recovery. Hence, the maintenance of the intensity of the flow alteration scheme by the numerous water withdrawals and the low tributary contributions, coupled with differential geomorphological characteristics along the reach, overwhelmed the natural tendency for the river to restore its natural conditions with distance downstream. Improving water management and, particularly, restoring endangered riparian ecosystems require a detailed understanding of existing and potential woody species behaviour across the geomorphological settings of rivers. Copyright © 2012 John Wiley & Sons, Ltd.     

TRACING GROUNDWATER DISCHARGE IN THE FLOODPLAIN OF THE PARANA RIVER,ARGENTINA: IMPLICATIONS FOR ITS BIOLOGICAL COMMUNITIES

Groundwater discharge can be an important determinant of the functioning of aquatic environments and their associated biological communities. However, the presence and the importance of groundwater have not been considered in the Parana River floodplain owing to the large quantity of surface water. The present study aimed to identify groundwater discharge conditions in a sector of the middle of the floodplain of the Parana River by studying groundwater flow systems. Eight piezometers were installed to record groundwater movement in the vertical plane weekly for 2 years. Water samples were collected in piezometers, domestic wells, the river and other water bodies to study the groundwater flow systems. Rising of the water level during piezometer installation and recording suggested that the study zone represents groundwater discharge conditions. Residence time proxy allowed identification of local flows and intermediate flows. Local rainfall (in Santa Fe) showed an isotopic signature (δD and δ¹⁸O) similar to some local flows detected in the study zone, and this suggested local recharge. The chemical characteristics of an intermediate flow suggest that water would have travelled from a recharge area ~30 km from the study zone. Local rainfall and the intermediate flow have different isotopic signature. Results suggest that the willow forest is associated with the recharge area of a local flow, which plurispecific‐canopy forest is related to a transit area of a semi‐intermediate flow, and that tall grassland and marshy community colonize discharge areas of local and intermediate flows, respectively. Copyright © 2012 John Wiley & Sons, Ltd.     

EFFECTS OF FLOW DYNAMICS ON THE AQUATIC‐TERRESTRIAL TRANSITION ZONE (ATTZ) OF LOWER MISSOURI RIVER SANDBARS WITH IMPLICATIONS FOR SELECTED BIOTA

Sandbars are an important aquatic terrestrial transition zone (ATTZ) in the active channel of rivers that provide a variety of habitat conditions for riverine biota. Channelization and flow regulation in many large rivers have diminished sandbar habitats and their rehabilitation is a priority. We developed sandbar‐specific models of discharge‐area relationships to determine how changes in flow regime affect the area of different habitat types within the submerged sandbar ATTZ (depth) and exposed sandbar ATTZ (elevation) for a representative sample of Lower Missouri River sandbars. We defined six different structural habitat types within the sandbar ATTZ based on depth or exposed elevation ranges that are important to different biota during at least part of their annual cycle for either survival or reproduction. Scenarios included the modelled natural flow regime, current managed flow regime and two environmental flow options, all modelled within the contemporary river active channel. Thirteen point and wing‐dike sandbars were evaluated under four different flow scenarios to explore the effects of flow regime on seasonal habitat availability for foraging of migratory shorebirds and wading birds, nesting of softshell turtles and nursery of riverine fishes. Managed flows provided more foraging habitat for shorebirds and wading birds and more nursery habitat for riverine fishes within the channelized reach sandbar ATTZ than the natural flow regime or modelled environmental flows. Reduced summer flows occurring under natural and environmental flow alternatives increased exposed sandbar nesting habitat for softshell turtle hatchling emergence. Results reveal how management of channelized and flow regulated large rivers could benefit from a modelling framework that couples hydrologic and geomorphic characteristics to predict habitat conditions for a variety of biota. Published in 2011 by John Wiley & Sons, Ltd.