CLASSIFICATION OF FLOW REGIMES FOR ENVIRONMENTAL FLOW ASSESSMENT IN REGULATED RIVERS: THE HUAI RIVER BASIN,CHINA

Flow regime characteristics (magnitude, frequency, duration, seasonal timing and rates of change) play a primary role in regulating the biodiversity and ecological processes in rivers. River classification provides the foundation for comparing the hydrologic regimes of rivers and development of hydro‐ecological relationships to inform environmental flow management and river restoration. This paper presents a classification of natural flow regimes and hydrologic changes due to dams and floodgates in the Huai River Basin, China, in preparation for an environmental flow assessment. The monthly natural flow regime of 45 stations in the upper and middle Huai River Basin were simulated for the period 1963–2000, based on the hydrological model SWAT (Soil and Water Assessment Tool). Six classes of flow patterns (low or high discharge, stable or variable, perennial or intermittent, predictable or unpredictable) were identified based on 80 hydrologic metrics, analysed by hierarchical clustering algorithms. The ecologically relevant climatic and geographic characteristics of these flow classes were tested for concordance with, and to strengthen, the hydro‐ecological classification. The regulation of natural flow patterns by dams and floodgates changed flows at some locations within each flow class and caused some gauges to shift into another class. The research reported here is expected to provide a foundation for development of hydro‐ecological relationships and environmental flow methods for wider use in China, as well as setting a new scientific direction for integrated river basin management in the Huai River Basin. Copyright © 2011 John Wiley & Sons, Ltd.     

Hydrological changes and ecological impacts associated with water resource development in large floodplain rivers in the Australian tropics

The majority of rivers in the Australian tropics possess near‐natural flow regimes and are an ecological asset of global significance. We examined flow variability in large floodplain rivers in the Gulf of Carpentaria, northern Australia, and the potential ecological impacts of future water resource development (WRD). Flow metrics based on long‐term records were used to classify flow regimes and predict hydrological drivers of ecological function. Flow regimes of selected rivers were then compared with those simulated for pre‐ and post‐WRD flows in the Darling River, a highly modified river in Australia's south‐east. Generally, rivers were classified as typically ‘tropical’ (more permanent, regular flows) or ‘dryland’ (more ephemeral, with greater flow variability). In addition, all rivers displayed wet–dry seasonality associated with changes in flow magnitude or number of zero‐flow days. We propose that these features (flow permanence and regularity; flow variability and absence; wet–dry seasonality) are the key hydrological drivers of biodiversity and ecological function in the floodplain rivers of Australia's north. In terms of WRD, inter‐annual flow variability was predicted to increase or decrease depending on rivers' natural flow regimes, specifically their tendency toward lower or higher flow magnitudes. Either outcome is expected to have adverse effects on the biodiversity and ecological function of these relatively pristine rivers and floodplain habitats. In particular, reduced and homogenized habitat, loss of life‐history cues, inhibited dispersal and shifts in community composition, as a result of WRD, threaten the ecological integrity of rivers adapted to the three hydrological drivers above. These findings serve as a caution for careful consideration of WRD options for rivers in the Australian tropics and for those with similar flow regimes the world over. Copyright © 2008 John Wiley & Sons, Ltd.     

HYDROGEOMORPHIC CLASSIFICATION OF WASHINGTON STATE RIVERS TO SUPPORT EMERGING ENVIRONMENTAL FLOW MANAGEMENT STRATEGIES

As demand for fresh water increases in tandem with human population growth and a changing climate, the need to understand the ecological tradeoffs of flow regulation gains greater importance. Environmental classification is a first step towards quantifying these tradeoffs by creating the framework necessary for analysing the effects of flow variability on riverine biota. Our study presents a spatially explicit hydrogeomorphic classification of streams and rivers in Washington State, USA and investigates how projected climate change is likely to affect flow regimes in the future. We calculated 99 hydrologic metrics from 15 years of continuous daily discharge data for 64 gauges with negligible upstream impact, which were entered into a Bayesian mixture model to classify flow regimes into seven major classes described by their dominant flow source as follows: groundwater (GW), rainfall (RF), rain‐with‐snow (RS), snow‐and‐rain (SandR), snow‐with‐rain (SR), snowmelt (SM) and ultra‐snowmelt (US). The largest class sizes were represented by the transitional RS and SandR classes (14 and 12 gauges, respectively), which are ubiquitous in temperate, mountainous landscapes found in Washington. We used a recursive partitioning algorithm and random forests to predict flow class based on a suite of environmental and climate variables. Overall classification success was 75%, and the model was used to predict normative flow classes at the reach scale for the entire state. Application of future climate change scenarios to the model inputs indicated shifts of varying magnitude from snow‐dominated to rain‐dominated flow classes. Lastly, a geomorphic classification was developed using a digital elevation model (DEM) and climatic data to assign stream segments as either dominantly able or unable to migrate, which was cross‐tabulated with the flow types to produce a 14‐tier hydrogeomorphic classification. The hydrogeomorphic classification provides a framework upon which empirical flow alteration–ecological response relationships can subsequently be developed using ecological information collected throughout the region. Copyright © 2011 John Wiley & Sons, Ltd.     

River flow indexing using British benthic macroinvertebrates: a framework for setting hydroecological objectives

A method linking qualitative and semi‐quantitative change in riverine benthic macroinvertebrate communities to prevailing flow regimes is proposed. The Lotic‐invertebrate Index for Flow Evaluation (LIFE) technique is based on data derived from established survey methods, that incorporate sampling strategies considered highly appropriate for assessing the impact of variable flows on benthic populations. Hydroecological links have been investigated in a number of English rivers, after correlating LIFE scores obtained over a number of years with several hundred different flow variables. This process identifies the most significant relationships between flow and LIFE which, in turn, enables those features of flow that are of critical importance in influencing community structure in different rivers to be defined. Summer flow variables are thus highlighted as being most influential in predicting community structure in most chalk and limestone streams, whereas invertebrate communities colonizing rivers draining impermeable catchments are much more influenced by short‐term hydrological events. Biota present in rivers with regulated or augmented flows tend to be most strongly affected by non‐seasonal, interannual flow variation. These responses provide opportunities for analysing and elucidating hydroecological relationships in some detail, and it should ultimately be possible to use these data to set highly relevant, cost‐effective hydroecological objectives. An example is presented to show how this might be accomplished. Key areas of further work include the need to provide robust procedures for setting hydroecological objectives, investigation of habitat quality and LIFE score relationships in natural and degraded river reaches and evaluation of potential links with other biological modelling methods such as RIVPACS. Copyright © 1999 John Wiley & Sons, Ltd.     

Combined Effects of Reservoir Operations and Climate Warming on the Flow Regime of Hydropower Bypass Reaches of California's Sierra Nevada

Alterations to flow regimes from regulation and climatic change both affect the biophysical functioning of rivers over long time periods and large spatial areas. Historically, however, the effects of these flow alteration drivers have been studied separately. In this study, results from unregulated and regulated river management models were assessed to understand how flow regime alterations from river regulation differ under future climate conditions in the Sierra Nevada of California, USA. Four representative flow alteration metrics—mean annual flow, low flow duration, centroid timing and mean weekly rate of decrease—were calculated and statistically characterized under historical and future unregulated and regulated conditions over a 20‐year period at each of the eight regulated river locations below dams across the Sierra Nevada. Future climatic conditions were represented by assuming an increase in air temperature of 6 °C above historical (1981–2000) air temperatures, with no change in other meteorological conditions. Results indicate that climate warming will measurably alter some aspects of the flow regime. By comparison, however, river regulation with business‐as‐usual operations will alter flow regimes much more than climate warming. Existing reservoirs can possibly be used to dampen the anticipated effects of climate warming through improved operations, though additional research is needed to identify the full suite of such possibilities. Copyright © 2014 John Wiley & Sons, Ltd.     

Macroinvertebrate community response to inter‐annual and regional river flow regime dynamics

Spatio‐temporal variability in river flow is a fundamental control on instream habitat structure and riverine ecosystem biodiversity and integrity. However, long‐term riverine ecological time‐series to test hypotheses about hydrology–ecology interactions in a broader temporal context are rare, and studies spanning multiple rivers are often limited in their temporal coverage to less than five years. To address this research gap, a unique spatio‐temporal hydroecological analysis was conducted of long‐term instream ecological responses (1990–2000) to river flow regime variability at 83 sites across England and Wales. The results demonstrate clear hydroecological associations at the national scale (all data). In addition, significant differences in ecological response are recorded between three ‘regions’ identified (RM1–3*) associated with characteristics of the flow regime. The effect of two major supra‐seasonal droughts (1990–1992 and 1996–1997) on inter‐annual (IA) variability of the LIFE scores is evident with both events showing a gradual decline before and recovery of LIFE scores after the low flow period. The instream community response to high magnitude flow regimes (1994 and 1995) is also apparent, although these associations are less striking. The results demonstrate classification of rivers into flow regime regions offers a way to help unravel complex hydroecological associations. The approach adopted herein could easily be adapted for other geographical locations, where datasets are available. Such work is imperative to understand flow regime–ecology interactions in a longer term, wider spatial context and so assess future hydroecological responses to climate change and anthropogenic modification of riverine ecosystems. Copyright © 2008 John Wiley & Sons, Ltd.     

Potential Effects of Climate Change on Ecologically Relevant Streamflow Regimes

We assessed the climate‐driven changes in ecologically relevant flow regimes expected to occur by the year 2100 in streams across the conterminous United States. We used long‐term daily flow measurements from 601 gauged streams whose watersheds were in relatively natural condition to characterize spatial variation in 16 flow regime variables selected for their ecological importance. Principal component analysis of these 16 variables produced five uncorrelated factors that described patterns of spatial covariation in flow regimes. These five factors were associated with low flow, magnitude, flashiness, timing, and constancy characteristics of the daily flow regime. We applied hierarchical clustering to the five flow factors to classify the 601 streams into three coarses and eight more finely resolved flow regime classes. We then developed a random forest model that used watershed and climate attributes to predict the probabilities that streams belonged to each of the eight finely resolved flow regime classes. The model had a prediction accuracy (per cent correct classification) of 75%. We used the random forest model with downscaled climate (precipitation and temperature) projections to predict site‐specific changes in flow regime classes expected by 2100. Thirty‐three per cent of the 601 sites were predicted to change to a different flow regime class by 2100. Snow‐fed streams in the western USA were predicted to be less likely to change regimes, whereas both small, perennial, rain‐fed streams and intermittent streams in the central and eastern USA were predicted to be most likely to change regime. Copyright © 2016 John Wiley & Sons, Ltd.     

CLASSIFICATION OF NATURAL FLOW REGIMES IN THE EBRO BASIN (SPAIN) BY USING A WIDE RANGE OF HYDROLOGIC PARAMETERS

This paper presents a classification of different natural flow regimes found in Ebro basin, one of the largest in the Mediterranean region. Determination of flow regimes was based on multivariate analyses using long‐term discharge series of unaltered flow data. Mean monthly discharges of the 30 ‘best’ flow series and a total of 52 flow series containing unaltered flow data were selected to represent baseline flow conditions for tributaries throughout the basin. Metrics representing magnitude, duration and frequency components of flow were used to identify hydrologic differences across the basin. A total of six natural flow regimes were identified in the Ebro Basin, using a Ward cluster method. The flow patterns identified and their spatial distribution largely corresponded with climatic zones previously reported for the Ebro Basin, with regime types ranging from pluvio‐oceanic in the western part of the basin to Mediterranean in the eastern region. Geologic characteristics of the catchment and altitude of headwaters were also found to play an important role in defining flow regime type. A 19‐hydrologic variable subset was used to explain main hydrologic differences among groups (such as magnitude and frequency of extreme flow conditions or magnitude and variance of average flow conditions). However, stepwise discriminant analysis was not able to identify consistent subsets of hydrologic variables that adequately identified the six natural flow regime types in this basin. Canonical discriminant analysis was useful to understand class separation and for the interpretation of results. Copyright © 2012 John Wiley & Sons, Ltd.     

How freshwater biomonitoring tools vary sub‐seasonally reflects temporary river flow regimes

Characterizing temporary river ecosystem responses to flow regimes is vital for conserving their biodiversity and the services they provide to society. However, freshwater biomonitoring tools rarely reflect community responses to hydrological variations or flow cessation events, and those available have not been widely tested within temporary rivers. This study examines two invertebrate biomonitoring tools characterizing community responses to different flow‐related properties: the “Drought Effect of Habitat Loss on Invertebrates” (DEHLI) and “Lotic‐invertebrate Index for Flow Evaluation” (LIFE), which, respectively reflect community responses to habitat and hydraulic properties associated with changing flow conditions. Sub‐seasonal (monthly) variations of LIFE and DEHLI were explored within two groundwater‐fed intermittent rivers, one dries sporadically (a flashy, karstic hydrology—River Lathkill) and the other dries seasonally (a highly buffered flow regime—South Winterbourne). Biomonitoring tools were highly sensitive to channel drying and also responded to reduced discharges in permanently flowing reaches. Biomonitoring tools captured ecological recovery patterns in the Lathkill following a supra‐seasonal drought. Some unexpected results were observed in the South Winterbourne where LIFE and DEHLI indicated relatively high‐flow conditions despite low discharges occurring during some summer months. This probably reflected macrophyte encroachment, which benefitted certain invertebrates (e.g., marginal‐dwelling taxa) and highlights the importance of considering instream habitat conditions when interpreting flow regime influences on biomonitoring tools. Although LIFE and DEHLI were positively correlated, the latter responded more clearly to drying events, highlighting that communities respond strongly to the disconnection of instream habitats as flows recede. The results highlighted short‐term ecological responses to hydrological variations and the value in adopting sub‐seasonal sampling strategies within temporary rivers. Findings from this study indicate the importance of establishing flow response guilds which group taxa that respond comparably to flow cessation events. Such information could be adopted within biomonitoring practices to better characterize temporary river ecosystem responses to hydrological variations.     

Classification of reaches in the Missouri and lower Yellowstone Rivers based on flow characteristics

Several aspects of flow have been shown to be important determinants of biological community structure and function in streams, yet direct application of this approach to large rivers has been limited. Using a multivariate approach, we grouped flow gauges into hydrologically similar units in the Missouri and lower Yellowstone Rivers and developed a model based on flow variability parameters that could be used to test hypotheses about the role of flow in determining aquatic community structure. This model could also be used for future comparisons as the hydrological regime changes. A suite of hydrological parameters for the recent, post‐impoundment period (1 October 1966–30 September 1996) for each of 15 gauges along the Missouri and lower Yellowstone Rivers were initially used. Preliminary graphical exploration identified five variables for use in further multivariate analyses. Six hydrologically distinct units composed of gauges exhibiting similar flow characteristics were then identified using cluster analysis. Discriminant analyses identified the three most influential variables as flow per unit drainage area, coefficient of variation of mean annual flow, and flow constancy. One surprising result was the relative similarity of flow regimes between the two uppermost and three lowermost gauges, despite large differences in magnitude of flow and separation by roughly 3000 km. Our results synthesize, simplify and interpret the complex changes in flow occurring along the Missouri and lower Yellowstone Rivers, and provide an objective grouping for future tests of how these changes may affect biological communities. Copyright © 2002 John Wiley & Sons, Ltd.     

Hydrological classification of non-perennial Mediterranean rivers and streams: A new insight for their management within the water framework directive

Classification of natural flow regimes of non-perennial rivers and streams (NPRS) is an incipient field of research. NPRS represent approximately 70% of the total Mediterranean rivers and are expected to increase in the next decades as a result of climate change. Due to the ecological importance of NPRS and the need to improve national ecological assessment methods within the scope of the Water Framework Directive (WFD), this paper aims to classify the hydrological regime of 69 non-regulated streams, testing several hydrological indices related to the magnitude, frequency, duration, timing, and rate of change in periods of flow cessation. Using daily flow records, a total of 315 indices were calculated and their relationships were examined with Principal Component Analysis (PCA) for different thresholds used to define zero-flow condition (0, 1, 2, and 5 L/s). Redundancy analysis identified five indices that better describe the patterns of hydrological variability in Mediterranean NPRS: number of days per year without flow, annual percentage of months without flow, mean daily annual flow, coefficient of variation of Julian date of the annual start of zero-flow and annual rise rate. Using these indices, a self-organizing map (SOM) was trained in order to categorize the NPRS into three groups with similar hydrological features. The results of this study provide a statistically-based hydrological classification of NPRS in Mediterranean environments. We expect that this classification will provide useful insights to water authorities to improve the assessments of the ecological status in this type of water bodies.     

The Hydrological Status Concept: Application at a Temporary River (Candelaro,Italy)

In achieving the final objective of the European Water Framework Directive, the evaluation of the ‘hydrological status’ of a water body in a catchment is of the utmost importance. It represents the divergence of the actual hydrological regime from its ‘natural’ condition and may thus provide crucial information about the ecological status of a river. In this paper, a new approach in evaluating the hydrological status of a temporary river was tested. The flow regime of a river has been classified through the analysis of two metrics: the permanence of flow and the predictability of no‐flow conditions that were evaluated on monthly streamflow data. This method was applied to the Candelaro river basin (Puglia, Italy) where we had to face the problem of limited data availability. The Soil and Water Assessment Tool model was used when streamflow data were not available, and a geographic information system procedure was applied to estimate potential water abstractions from the river. Four types of rivers were identified whose regimes may exert a control on aquatic life. By using the two metrics as coordinates in a plot, a graphic representation of the regime can be visualized in a point. Hydrological perturbations associated with water abstractions, point discharges and the presence of a reservoir were assessed by comparing the position of the two points representing the regime before and after the impacts. The method is intended to be used with biological metrics in order to define the ecological status of a stream, and it could also be used in planning the ‘measures’ aimed at fulfilling the Water Framework Directive goals. Copyright © 2014 John Wiley & Sons, Ltd.     

Macroinvertebrate Community Responses to Annual Flow Variation from River Regulation: An 11‐Year Study

The majority of the world's large river systems is affected by dams. The influences of unnatural regimes induced by flow management are wide‐ranging from both biotic and abiotic standpoints. However, many of these effects are not evident over short (1–2 years) periods (e.g. impacts of annual flow variation). This study examines the long‐term effects of annual flow variation on the macroinvertebrate community in the Chattahoochee River (GA) in the reaches below Buford Dam, the major water control structure on the river. Quarterly, macroinvertebrate samples were taken from 2001 to 2011 using Surber and Hester–Dendy plate samplers at six locations spread across 65 km below the dam. Data were analysed via analysis of similarities to determine differences in community composition between high‐flow (mean discharge = 58.27 m³/s) and low‐flow (mean discharge = 26.53 m³/s) years. Taxa that contributed most to community differences were determined via similarity percentages analyses and subsequent t‐tests. Several insect taxa (e.g. Cheumatopsyche and Ceratopsyche caddisfly larvae, Maccaffertium mayfly nymphs and Taeniopteryx stonefly nymphs) were more prevalent under the high‐flow regime. Non‐insect macroinvertebrates (e.g. Crangonyx amphipods, Tricladida flatworms and Caecidotea isopods) were more abundant under low‐flow conditions. In terms of taxon richness, no significant effects of flow regime were detected. Implications of macroinvertebrate patterns for the fishery and ecological health of the river are discussed. Copyright © 2014 John Wiley & Sons, Ltd.     

Comparative analysis of glacial and nival streamflow regimes with implications for lotic habitat quantity and fish species richness

Growing interest in the differential responses of glacial and nival rivers to climatic forcing, and in ecological distinctions between the two streamflow regimes, suggests the need for a better comparative understanding of how the annual hydrologic cycle differs with presence or absence of catchment glacial cover. In this study, timing and magnitude characteristics of the average annual hydrographs of five glacierized and four nival catchments in the southwestern Canadian subarctic are empirically identified and compared. Likely effects upon fish habitat are qualitatively assessed, and net fisheries potential is tentatively investigated using taxa richness data. The chief hydrological conclusions at P < 0.05 using Kolmogorov–Smirnov and empirical orthogonal function analyses are: (1) catchment glacial cover results in freshets that are longer, larger, and peak later than those experienced by the nival regime; (2) the winter baseflows of glacial rivers are also much higher on a unit‐catchment‐area basis; and (3) basin scale and degree of catchment glacial cover are of comparable importance in determining the magnitude of the annual hydrologic cycle. These differences arise from the greater availability, both in volume and over time, of meltwater in glacial catchments, which in part reflects the consistently negative alpine glacial mass balances observed both in the present study area and globally under historical climatic warming. Such regime distinctions result in increased spawning season and winter aquatic habitat availability, which may in turn offset negative habitat characteristics previously identified for glacial river ecosystems. While previous studies have suggested that glacial influences tend to decrease macroinvertebrate diversity and increase salmon populations, preliminary analysis of available fish species presence/absence data from the current study area tentatively appears to suggest similar or, perhaps, slightly higher fish taxa richness relative to nival streams; in all three cases, however, catchment lake cover may play a key hydroecological modifying role. The results strongly confirm and extend existing understanding of glacial–nival regime differences with respect to both streamflow and fisheries ecology, and raise new questions for future research. Copyright © 2005 John Wiley & Sons, Ltd.     

NATURAL FLOW REGIME CLASSIFICATIONS ARE SENSITIVE TO DEFINITION PROCEDURES

The correspondence and performance of six classifications of flow regimes of New Zealand rivers that were all mapped onto the same digital river network were assessed. Classification 1 was defined deductively, based on expert‐defined rules. Classifications 2 to 6 were defined inductively using hydrological indices calculated from 321 natural daily flow records. Classifications 2 to 4 were defined by first clustering the gauges based on the hydrological indices and then predicting the class of each segment of the network using a Random Forest classifier. Classifications 5 and 6 were defined by first predicting the indices for each segment of the network using Random Forest regression models. Cluster analysis was then used to group the network segments into classes. Further differences between classifications were due to differences in the standardisation of the hydrological indices and clustering algorithms. Correspondence (extent to which the patterns defined by the classifications were similar) was assessed formally using the adjusted Rand index and visually. The performance of the classifications was assessed using classification strength calculated using the hydrological indices and ANOVA calculated for individual indices. Correspondence between the classifications was low (adjusted Rand index range, 0.1–0.5). Classification strength and ANOVA statistics assessed using cross validation indicated that the inductive classifications performed better than the deductively defined classification and that there were some significant differences in performance between the inductive classifications. However, these differences were not large from a practical point of view. Our results indicate that there are many credible classifications of the flow regimes of a study region. When considering methods for defining flow regime classifications, aspects other than the predictive performance, such as flow data requirements and how easily the final classification can be explained, should be considered. Copyright © 2012 John Wiley & Sons, Ltd.     

Classification of natural flow regimes in Poland

Hydrological classifications are aimed at simplifying spatiotemporal variability of flow regimes and, secondly, at supporting environmental flow management. The objective of this study was to perform classification of natural flow regimes in Poland using an inductive approach based on a set of hydrological metrics (HMs) and to develop a model for prediction of class membership based on a set of environmental variables (EVs). A set of 147 gauges with unmodified flow regimes was identified, and for each gauge, values of 73 HMs and 28 EVs were computed. Classification was performed using k‐means and k‐medoids techniques, based on 4 principal components explaining 73.4% of variability in HMs. Out of 7 distinguished classes, 4 (P1–P4) were spread across the Polish Plain, 1 (U5) was restricted to uplands, and 2 (M6 and M7) to mountains. The between‐class differences in HMs and EVs were generally high, although classes P1 and P3 were not easily distinguishable. Mean predictive accuracy of the developed random forest model was 79%, which is high compared to other studies of this type. The lowest accuracies (0 and 50%) were achieved by 2 classes with the lowest counts. Variables representing diverse aspects: hydrography, climate, topography, and geology had the highest importance in the random forest model. Future research can benefit from the database of selected gauges with computed HMs, EVs, and assigned classes, freely available through a long‐lasting data repository. With this study, the first step towards application of the ELOHA framework for environmental flow management at regional scale has been achieved.     

Climate and rivers

Over the last few decades as hydrologists have slowly raised their line of sight above the watershed boundary, it has become increasingly recognised that what happens in the atmosphere, as a major source of moisture for the terrestrial branch of the hydrological cycle, can strongly influence river dynamics at a range of spatial and temporal scales. Notwithstanding this, there is still a tendency for some in the river research community to restrict their gaze to the river channel or floodplain. However, Geoff Petts, the person to which this special issue is dedicated, understood well and widely encouraged a holistic view of river catchment processes. This included an acknowledgment of the role of climate, in its broadest sense, in shaping what happens within and without the river channel. The purpose of this paper therefore is to offer a broad overview of the role of some aspects of climate science in advancing knowledge in river research. Topics to be addressed include the role of climate in influencing river flow regimes, a consideration of the large‐scale climate mechanisms that drive hydrological variability within river basins at interannual to decadal timescales and atmospheric rivers and their link to surface hydrology. In reviewing these topics, a number of key knowledge gaps have emerged including attributing the causes of river flow regime changes to any one particular cause, the nonstationary and asymmetric forcing of river regimes by modes of climate variability and establishing links between atmospheric rivers, and terrestrial river channel processes, fluvial habitats, and ecological change.     

Influence of Flow on Community Structure and Production of Snag‐Dwelling Macroinvertebrates in an Impaired Low‐Gradient River

The natural flow regime of rivers has been altered throughout the world in a variety of ways, with many alterations resulting in reduced flows. While restoring impaired systems remains a societal imperative, a fundamental understanding of the effects of reduced flows on river ecosystem structure and function is needed to refine restoration goals and guide implementation. We quantified the effects of chronic low flows on snag‐dwelling macroinvertebrate community structure and production in a low‐gradient river. Macroinvertebrates commonly associated with flowing water (e.g. passive filter‐feeders (PFF)) and higher quality habitats (e.g. Ephemeroptera, Plecoptera and Trichoptera (EPT)) had significantly higher abundance and biomass, and showed trends of higher production, in faster flowing reaches upstream of a hydrologic disconnection created by a drainage ditch. The presence of EPT and PFF groups resulted in a significantly more diverse community composed of larger‐sized individuals compared with downstream, low‐flow reaches, where smaller‐bodied taxa (e.g. small crustaceans), and groups reflective of degraded conditions (e.g. Oligochaeta, Isopoda and Chironomidae) dominated production. Multivariate analyses suggested that differences between these two disparate communities were driven by water velocity and organic matter resources. Mean estimates of total community production did not differ significantly between the two reaches, however, there were areas in low‐flow reaches that attained high secondary production because of patchily distributed and highly productive chironomids. Results demonstrate that long‐term reductions in flows, even in a low‐gradient river, can lead to significant shifts in macroinvertebrate communities, ultimately influencing energy flow pathways in stream food webs. Copyright © 2015 John Wiley & Sons, Ltd.