Seed banks and seed survival of submersion for an emerging plant invader in the Colorado River system,USA

Soil seed banks along shorelines and the ability of seeds to survive submersion in water are among potential factors that can influence seed supply, a frequent driver of riparian plant invasions. We investigated germinable soil seed banks and the ability of seeds to germinate after different durations of submersion in water for non-native ravennagrass (Saccharum ravennae), an emerging but poorly understood invader of riparian habitats in the American Southwest, including along the Lake Powell section of the Colorado River where we performed the study. We assessed soil seed bank composition in 27 ravennagrass-invaded plots using the emergence technique and evaluated effects on ravennagrass germination of submersion in river water for up to 15 months in a laboratory experiment. Ravennagrass was sparse in soil seed banks, detected in samples from only one of the 27 plots. Instead, soil seed banks were dominated by the non-native annual cheatgrass (Bromus tectorum) and a mixture of native early colonizers (e.g., Pseudognaphalium stramineum) and shrubs (e.g., Baccharis emoryi). Although ravennagrass seed germination was low (<9%) in the submersion experiment, some seeds retained germinability after 15 months of continuous submersion in water. Results suggest that while ravennagrass may have limited on-site, readily germinable soil seed banks, potential implications of its protracted seed viability in water for dispersal along waterways warrants further attention.     

Secondary seed dispersal in hydro‐fluctuation belts and its influence on the soil seed bank

The completion of the Three Gorges project altered the water level fluctuations from “summer submersion and winter exposure” to “summer exposure and winter submersion.” The reverse seasonal flooding has dramatically changed both the time and duration of exposure and submergence of reservoir banks. Secondary seed dispersal by water is the primary method of dispersal in hydro‐fluctuation belts. This study conducted field sampling and germination experiments to evaluate the effects of reverse seasonal flooding on species composition, diversity, and density of the secondary seed dispersal system and the soil seed bank, and their changes with elevation, in the Xiangzi River. The results indicate that that the secondary seed dispersal system and soil seed bank were associated with 42 and 50 plant species, respectively, and their average seed densities were 1,876.27 and 7,322.18 seeds/m², respectively. The quantity of seeds from secondary seed dispersal accounted for 20.40% of the soil seed bank. The species life‐form composition of both consisted mainly of annual and perennial grasses. Among these grasses, the dominant species in the secondary seed dispersal system were Cynodon dactylon and Paspalum distichum, whereas the dominant species in the soil seed bank were Solanum nigrum and Rumex dentatus. Due to the effects of reverse seasonal water level fluctuations, the species composition and quantity of the soil seed bank and secondary seed dispersal system largely exhibited similar spatial distribution characteristics, but some differences were observed. The species quantity, species diversity index, and seed density in the soil seed bank were the highest in the middle section of the hydro‐fluctuation belt, followed by the top section and then the bottom section. Conversely, the species quantity, species diversity index, and seed density in the secondary seed dispersal system were highest at the top section, followed by the middle section and then the bottom section.     

鄱阳湖区堤防植物多样性特征与土壤种子库密度的关系

以鄱阳湖区重点堤防作为研究对象,采用线路踏查、样方调查和种子萌发相结合的方法,研究不同加固土料堤防地上植物多样性特征与土壤种子库密度的关系。结果表明:湖区重点堤防高等植物共计106种,隶属于38科,89属,以草本植物为主,禾本科、菊科和大戟科为优势科;不同加固土料堤防植物多样性指数与土壤种子库密度以水稻土最高,其次是建筑垃圾,山地黏土最低;不同植草模式下0~10 cm土层的土壤种子库密度以自然修复模式最大,其次是假俭草植草模式,以狗牙根植草模式最小;土壤种子库密度与地上植物种数、Simpson指数、Shannon-weiner指数和Pielou群落均匀度指数呈显著正相关,与地上植被密度、重要值呈极显著正相关,但与Margalef丰富度指数无显著相关性。因此,堤防土壤种子库密度与地上植物多样性特征之间具有较强的相互作用关系,研究成果对于鄱阳湖区重点堤防的除险加固与管理维护具有重要意义。     

Can the Regeneration of Vegetation from Riparian Seed Banks Support Biogeomorphic Succession and the Geomorphic Recovery of Degraded River Channels?

For rivers degraded by erosion and channel widening, the re‐establishment of riparian vegetation is essential. We assess the potential for riparian seed banks to facilitate natural channel contraction through the regeneration of plants involved in the biogeomorphic succession of three discrete geomorphic units of increasing age and height above the channel bed: bars, benches and floodplain. Standing vegetation upon each unit type was surveyed for four river reaches in the Hunter catchment of eastern Australia. Seed bank composition was determined using seedling emergence techniques on sediment sampled from the units. We compared species richness and composition, and longevity, growth form and seed dispersal mechanisms between the standing vegetation and seed bank species. The seed bank was similar across bars, benches and floodplain, containing mostly perennial pioneer herbs, sedges and rushes, dispersed by wind and hydrochory (water transport). While bar vegetation was similar to the seed bank, bench and floodplain vegetation included later successional species such as shrubs and trees, significantly more grasses and vines (benches: χ²_{5, N = 402} = 102.033, p < 0.001; floodplain: χ²_{5, N = 792} = 30.324, p < 0.001) and higher proportions of unassisted and animal‐dispersed seeds (benches: χ²_{5, N = 352} = 89.409, p < 0.001; floodplain: χ²_{5, N = 338} = 56.026, p < 0.001). The results suggest that seed banks may support early stages of biogeomorphic succession, via regeneration of pioneer plants. However, plants, such as shrubs and trees that are observed upon units of increasing age and height above the channel bed (i.e. benches and floodplain), are likely sourced from transient seeds produced by local vegetation, rather than seed banks. Copyright © 2014 John Wiley & Sons, Ltd.     

Identifying the natural flow regime and the relationship with riparian vegetation for two contrasting western Australian rivers

The natural flow regime and the relationship between flows and riparian vegetation are described for sites on both the Blackwood River in south‐western Australia and the Ord River in north‐western Australia. Analysis of long‐term flow data showed the historic mean monthly river discharge for the Blackwood River is strongly seasonal and highly predictable with generally low variability each month. The Ord River showed a strong seasonality of flows with about 92% of the (total) yearly flow occurring between December and March. Flow variability was very high (e.g. coefficient of variation >100% for all months) but highly predictable, with this mostly attributed to low but constant dry‐season flows. Water depth, duration of flood events and the number of flood events per year show a significant correlation with aspects of the riparian vegetation within experimental vegetation plots. Results highlight the strong relationship between floristics, life form structure and population dynamics with stream hydrology. On the Blackwood River, species richness and cover of shrubs reduced with increased duration and frequency of flooding, while cover of exotic species and annual herbs increased with increased flooding. Germination of tree seedlings was not influenced by flood regime but size class of tree species increased with flooding frequency. On the Ord River, species richness was not influenced by flooding regime. However, cover of perennial grasses increased with flooding frequency whilst cover of shrubs decreased. There was no relationship between flooding and seedling establishment whilst tree size class decreased with increased flooding. The methods described here can be used to compare the response of different components of the riparian vegetation to different fluvial regimes (e.g. because of impoundment and abstraction). This technique can be expanded for the management of riparian zones and planning rehabilitation programmes. It may also be useful for improving the ecological knowledge base for setting environmental flows in regulated systems. Copyright © 2001 John Wiley & Sons, Ltd.     

Restoration of riparian vegetation on a mountain river degraded by historical mining and grazing

Riparian ecosystems in montane areas have been degraded by mining, streamflow alterations, and livestock grazing. Restoration of ecological and economic functions, especially in high-elevation watersheds that supply water to lower elevation urban and agriculture areas is of high priority. We investigated the response of riparian vegetation and bank stability following channel treatments and riparian habitat restoration along a segment of the upper Arkansas River south of Leadville, Colorado. The study area has been historically degraded by heavy-metal mining and is designated a U.S. Superfund site. Additionally, trans-basin water diversions and livestock grazing have contributed to channel widening and altered vegetation composition and cover. We used a before-after-control impact study design in four reaches with varied contamination and grazing history to assess restoration success. Before restoration, streambanks were dominated by graminoids and total vegetation cover varied among reaches with willow cover less than 16% in three reaches. Post-restoration, changes in total vegetation cover fell short of projected goals, but willow cover was greater than 20% in all study reaches. The increase in woody cover likely contributed to reduced erosion and vegetation encroachment post-restoration. Differences in functional group cover among reaches persisted post-restoration and may be attributed to soil contamination levels and low willow seed rain and dispersal. These results highlight the importance of setting realistic restoration goals based on elevation and past land use. We recommend further remediation of fluvial tailings with low vegetation cover and continued monitoring of willow height and cover to determine if further restoration activities are needed.     

Putting down roots: Afforestation and bank cohesion of Icelandic Rivers

Riparian vegetation is widely recognized as a critical component of functioning fluvial systems. Human pressures on woody vegetation including riparian areas have had lasting effects, especially at high latitude. In Iceland, prior to human settlement, native downy birch woodlands covered approximately 15%–40% of the land area compared to 1%–2% today. Afforestation efforts include planting seedlings, protecting native forest remnants, and acquiring land areas as national forests. The planted and protected nature of vegetation along rivers within forests provides a unique opportunity to evaluate the various taxa within riparian zones and the channel stabilizing characteristics of the vegetation used in afforestation. We investigated bank properties, sediment textures, and root characteristics within riparian zones along four rivers in forests in Iceland. Bank sediment textures are dominantly sandy loam overlying coarser textures. Undercut banks are common because of erosion of the less cohesive subsurface layer. Quantitative root data indicate that the woody taxa have greater root densities, rooting depths, and more complex root structures than forbs or graminoids. The native downy birch has the highest root densities, with <1 mm roots most abundant. Modeling of added bank cohesion indicates that willow provides up to six times and birch up to four times more added cohesion to the coarse sediment textures comprising stream banks compared to no vegetation. We conclude that planting and protecting the native birch and willow helps to reduce bank erosion, especially where long-term grazing exclusion can be maintained.     

Effects of stream flow intermittency on riparian vegetation of a semiarid region river (San Pedro River,Arizona)

The San Pedro River in the southwestern United States retains a natural flood regime and has several reaches with perennial stream flow and shallow ground water. However, much of the river flows intermittently. Urbanization‐linked declines in regional ground‐water levels have raised concerns over the future status of the riverine ecosystem in some parts of the river, while restoration‐linked decreases in agricultural ground‐water pumping are expected to increase stream flows in other parts. This study describes the response of the streamside herbaceous vegetation to changes in stream flow permanence. During the early summer dry season, streamside herbaceous cover and species richness declined continuously across spatial gradients of flow permanence, and composition shifted from hydric to mesic species at sites with more intermittent flow. Hydrologic threshold values were evident for one plant functional group: Schoenoplectus acutus, Juncus torreyi, and other hydric riparian plants declined sharply in cover with loss of perennial stream flow. In contrast, cover of mesic riparian perennials (including Cynodon dactylon, an introduced species) increased at sites with intermittent flow. Patterns of hydric and mesic riparian annuals varied by season: in the early summer dry season their cover declined continuously as flow became more intermittent, while in the late summer wet season their cover increased as the flow became more intermittent. Periodic drought at the intermittent sites may increase opportunities for establishment of these annuals during the monsoonal flood season. During the late summer flood season, stream flow was present at most sites, and fewer vegetation traits were correlated with flow permanence; cover and richness were correlated with other environmental factors including site elevation and substrate nitrate level and particle size. Although perennial‐flow and intermittent‐flow sites support different streamside plant communities, all of the plant functional groups are abundant at perennial‐flow sites when viewing the ecosystem at broader spatial and temporal scales: mesic riparian perennials are common in the floodplain zone adjacent to the river channel and late‐summer hydric and mesic annuals are periodically abundant after large floods. Copyright © 2005 John Wiley & Sons, Ltd.     

Resilience of riparian vegetation after restoration measures on River Inn

River restoration is widely applied, although its effects are poorly understood, and degraded habitats might be difficult to improve. Moreover, there is a lack of monitoring as well as few systematic comparisons of restoration methods. This study presents results of a 4‐year monitoring on River Inn (southern Germany) investigating restoration by gravel or sand addition or embankment removal. The results were compared with reference sites that represent the pre‐restoration conditions. At the landscape scale, we analysed vegetation types based on aerial photographs, whereas at a smaller scale, we undertook vegetation surveys and evaluated species composition, growth, and life form, as well as the proportion of the target vegetation. After 4 years, the data indicated a “negative resilience” of the vegetation back to the state prior to restoration. The structural analysis revealed an extensive spread of reed at expense of bare soil. Thus, the species composition largely regressed to the pre‐restoration conditions, and neither annuals nor other pioneer species showed a long‐term benefit of river restoration. There were differences among the three restoration treatments after 2 years, but no longer after 4 years. However, the river restoration had three positive outcomes: (a) There was a temporary benefit for pioneer vegetation that most likely replenished the seed bank of the respective species, (b) the valuable reed communities showed resilience, and (c) the measures allowed some practical learning as expected for adaptive restoration.     

Evidence for hydrochory and the deposition of viable seeds within winter flow‐deposited sediments: the River Dove,Derbyshire, UK

Previous studies of the influence of hydrochory on plant dispersal have focused on relationships with existing vegetation patterns or litter deposits. River sediment deposition studies have assessed sediment accretion rates, particle size and quality. The link between seed and sediment transport and deposition has been overlooked and never quantified. This paper presents observations of over‐winter viable seed and sediment deposition on three river margin sites along the River Dove, England. At these sites, 105 paired artificial turf mats were installed on the river bank top, face and toe from October 1999 to March 2000 to collect river‐deposited material. From each pair, one mat was used to determine the dry weight; median particle size; percentage sand, silt, clay and organic content of deposited sediment. The other was used in germination trials to determine the number and species of deposited viable seeds. Topographic surveys and water level measurements supported the estimation of the hydrological characteristics of each mat location. Few seeds and little sediment were found on mats not inundated by river water. For these inundated, sediment characteristics varied primarily with elevation (relative to local mean river water level during the study period), with less marked between‐site contrasts. The species composition of viable seeds also varied with elevation. Detrended correspondence analysis (DCA) revealed associations between river/wetland‐related seed species and the bank toe samples. Regression analysis revealed significant relationships between seed number, sediment properties and mat elevation. Redundancy analysis (RDA) revealed associations between the species and abundance of viable seeds and elevation, sediment weight and organic matter content. These analyses suggest that sediment and seed deposition by rivers are closely related processes. The implications of these results for seed remobilization and dispersal and hydrochory within riparian zones are discussed, as are the implications of changes in sediment/seed delivery mechanisms, river flow regimes, and the hydraulic properties of river margins for seeds, sediments and riparian vegetation patterns. Copyright © 2003 John Wiley & Sons, Ltd.     

Changes to herbaceous plant communities on a regulated desert river

Riparian plant communities are shaped by gradients of disturbance intensity and frequency and resource availability. Reservoir operation can alter the composition and abundance of riparian vegetation by changing the flood regime and by trapping fine sediments and associated nutrients within the reservoir system. We examined differences in herbaceous species richness, abundance and composition in Populus‐Salix stands along an unregulated and regulated reach of a river in semi‐arid Arizona, contrasted flood inundation frequency and edaphic conditions (soil moisture, nutrients and texture) between the reaches, and interpreted the vegetation differences in light of observed differences in environmental conditions. Flooding frequency was similar between reaches, but the proportion of fine textured soils in the unregulated reach was nearly double that of the regulated reach and soil nutrient levels were up to three times higher in the unregulated reach. Herbaceous cover and richness were consistently lower in the regulated reach, with between‐reach differences greatest during dry seasons. These patterns suggest that an edaphic‐based change in resource availability is the principal pathway by which river damming is altering herbaceous vegetation in this system. Our results demonstrate that sediment transport within riparian corridors is important for maintenance of herbaceous communities and that restoration of flow regimes alone may be insufficient to restore herbaceous flora on some regulated reaches. Copyright © 2008 John Wiley & Sons, Ltd.     

东江中游河边植被多样性调查评价

植被多样性作为河流健康评价体系的指标之一，反映了植被稳定性对河流健康程度的影响。通过对东江惠州段泗湄洲的植被进行调查分析，采用夏侬-威纳(Shannon-Weiner)多样性和均匀度评价法，对河边植被多样性进行了评价，分析得出了不同高程植被群落的物种组成。亚热带地域河边植被发育较快，植被演替主要决定于淹没频率与高程。从低滩到高滩，植被类型从草本向灌木、最后向木本过渡。植被的多样性也随着高程的增加而提高，高滩植被相对完整、稳定，其生物多样性指数比温带地区的指数高。水位的变化对低滩植被多样性的影响很大。低水位时，低滩散布多种草本植物；而高水位时，部分物种淹没死亡，余下以禾本科和沙草科植物为优势种草本植物群落。     

Vegetation development and restoration potential of drained reservoirs following dam removal in Wisconsin

Dam removal is potentially a powerful tool for river and riparian restoration. However, long‐term studies on the fate of former reservoirs do not exist, limiting assessment of the utility of dam removal as a means of riparian restoration. We took advantage of the decades‐long legacy of dam removals in Wisconsin to determine human uses of drained reservoirs and to evaluate vegetation establishment and species replacement at these sites. More than half of the 30 dam removal sites in southern Wisconsin over the past 47 years were used as commercial areas, parks and agricultural land, and active riparian restoration occurred on only two sites. For the 13 sites that were allowed to revegetate on their own, plants established in the first growing season and cover was very high at all sites in 2001. Species diversity and frequency (defined as percentage of sampled quadrats where a species is present) of trees were positively correlated with time since removal. No relationship existed between site age and frequencies of other growth forms, nor were there significant relationships between site age and the number or frequency of introduced species. However, mean frequency of introduced species was 75% per site and several sites were dominated by the introduced grass Phalaris arundinacea. Frequency of P. arundinacea was negatively correlated with number of native forbs, and lowest species diversity occurred on sites dominated by P. arundinacea. Ordination analyses revealed substantial site‐to‐site variation in vegetation that was weakly associated with gradients of site location, age, area, and soil phosphorus. Thus, temporal vegetation dynamics following dam removal were site‐specific. Rapid revegetation demonstrates the potential of these sites for riparian restoration. However, if dam removal is used as a means of restoring native riparian communities, then approaches must be tailored to individual sites and will need to focus on techniques to minimize establishment of aggressive invading species. Copyright © 2006 John Wiley & Sons, Ltd.     

Composition of the seed bank in drawdown areas of Navigation Pool 8 of the Upper Mississippi River

In an effort to enhance aquatic plant production and habitat diversity on the Upper Mississippi River (UMR), resource managers considered water level reduction as a management tool to increase the area of emergent and submersed aquatic vegetation by natural seed germination. To quantify the availability of seed, we assessed the potential seed bank of selected areas of Navigation Pool 8 of the UMR from substrate samples collected in spring 2000. We tested these samples for viable seed content under four hydrologic conditions: dry, moist, shallow flooded and submerged. Forty‐seven species were identified in the seed bank, including 27 obligate wetland, 10 facultative wetland and 7 upland species. Dominant taxa within the seed bank included Sagittaria spp., Lindernia dubia, Zosterella dubia, Cyperus spp., Eragrostis spp. and Leersia oryzoides. Of the four hydrologic treatments, moist substrates had the greatest species diversity and were the most productive, yielding an average density of 1420 seedlings m^?2. Emergent and submersed aquatic species were widely distributed, each type occurring in more than 90% of the samples. Timing of seedling germination varied among species and has implications for scheduling drawdowns to promote establishment of desired species. Seed bank results were correlated with the vegetation response on substrates exposed during a reduction of water levels of Pool 8 during summer 2001. Experimentally determining the composition and viability of seed banks from drawdown areas provides information useful in predicting the types of vegetation that may develop on exposed substrates. Further, these findings provide resource managers a better understanding of the potential for achieving desired vegetation response through water level reductions. Published in 2008 by John Wiley & Sons, Ltd.     

Sorting of seeds by hydrochory

The species composition of riparian vegetation is determined by a variety of processes, including the dispersal of seeds. The seed types that end up at certain locations are determined by their particular characteristics as well as the prevailing hydrology and the physical characteristics of the river channel. A conceptual model is proposed to identify the hydraulic processes involved in the hydrochory pathway that may lead to differential response of different seed types and hence their sorting. The results of a series of laboratory experiments are presented to demonstrate the sorting between seed types by these processes. The behaviour of non‐buoyant seeds during settling and entrainment is shown to be consistent with that of mineral sediments, and their sorting can therefore be described by existing sediment transport theory. Significant sorting of buoyant seed types takes place during transport under the influence of wind and during ingress into marginal emergent vegetation, trapping by vegetation and stranding during receding flows on sloping channel banks. Copyright © 2008 John Wiley & Sons, Ltd.     

Life‐History Plasticity of Riparian Annual Plants Adapted to Extreme Variations in Water Level: Mesocosm Experiments

The riparian zones of reservoirs associated with regulated rivers in China experience annual fluctuations in water level of up to 30 m that may vary in timing from year to year. Few plant species can tolerate such hydrological perturbation, but short‐lived riparian annuals might be evolutionarily pre‐adapted to such conditions. This study investigated plasticity of life history in four annual species: one typically associated with free‐flowing rivers (Panicum bisulcatum) and three that colonize reservoir margins (Cyperus michelianus, Fimbristylis miliacea and Eclipta prostrata). We found that all four species produced non‐dormant seeds that survived prolonged submergence; germination percentage was independent of the time of exposure by receding waters. Although growth was reduced as a result of shorter growing seasons, all four species completed their life cycles and produced seeds before winter. In addition, P. bisulcatum and C. michelianus allocated biomass to seed production, at the expense of roots and stems, in response to later establishment. All species responded to later establishment with a reduced vegetative growth period before seed production. C. michelianus, F. miliacea and E. prostrate could also delay the onset of flowering time by up to 2 months. P. bisulcatum, a plant that can flower only after exposure to short days, consequently had a fixed flowering time and could accommodate delayed establishment only with a progressively shorter period of vegetative growth. This lower flexibility might explain its absence from reservoir margins. The conceptual framework presented here offers a tool to predict the establishment of vegetation under hydrological disturbance in riparian environments and thereby provides insights into improved restoration practice. Copyright © 2014 John Wiley & Sons, Ltd.     

不同植被重建模式下赣北红壤侵蚀坡地土壤养分效应

为了探讨植被重建模式与土壤养分的关系,合理改良与利用红壤侵蚀坡地,本文分析了草本、果树、果树+草、果树+草+农作物和果树+农作物等不同植被重建模式对土壤养分的影响,并运用灰色关联分析方法对不同恢复模式的土壤养分效应进行综合评价。研究结果表明:不同植被重建模式土壤养分效应差异显著,以果树+横坡套种农作物改善土壤养分效应最好,然后依次是果树+草本模式(全区覆盖优于带状覆盖)、果+草+农作物模式、果树+纵坡套种农作物,草本、果树模式改善土壤养分效应最差。赣北侵蚀坡地植被恢复,可以优先考虑恢复果树+横坡套种农作物模式和果树+草本模式。     

Quantifying the effect of riparian forest versus agricultural vegetation on river meander migration rates,central Sacramento River,California, USA

Riparian forest vegetation is widely believed to protect riverbanks from erosion, but few studies have quantified the effect of riparian vegetation removal on rates of river channel migration. Measured historical changes in a river channel centreline, combined with mapped changes in floodplain vegetation, provide an opportunity to test how riparian vegetation cover affects the erodibility of riverbanks. We analysed meander migration patterns from 1896 to 1997 for the central reach of the Sacramento River between Red Bluff and Colusa, using channel planform and vegetation cover data compiled from maps and aerial photography. We used a numerical model of meander migration to back‐calculate local values for bank erodibility (i.e. the susceptibility of bank materials to erosion via lateral channel migration, normalized for variations in near‐bank flow velocities due to channel curvature). A comparison of migration rates for approximately 50 years before and after the construction of Shasta dam suggests that bank migration rates and erodibility increased roughly 50%, despite significant flow regulation, as riparian floodplains were progressively converted to agriculture. A comparison of migration rates and bank erodibilities between 1949 and 1997, for reaches bordered by riparian forest versus agriculture, shows that agricultural floodplains are 80 to 150% more erodible than riparian forest floodplains. An improved understanding of the effect of floodplain vegetation on river channel migration will aid efforts to predict future patterns of meander migration for different river management and restoration scenarios. Copyright © 2004 John Wiley & Sons, Ltd.     

Restoration of Hydrochory Following Dam Removal on the Elwha River,Washington

Hydrochory, seed dispersal by water, affects riparian vegetation by contributing to downstream community composition and diversity. However, dams can block hydrochory, reducing downstream species diversity and fragmenting riparian corridors. Dam removal is becoming more prevalent for economic and ecological reasons and is expected to restore hydrochory; however, this has never been documented in rivers. The largest dam removal project to date was the 2011 to 2014 removal of the Glines Canyon and Elwha dams on the Elwha River in Washington. Prior to dam removal, hydrochory was lower below Glines Canyon Dam compared with an upstream reach; our objective was to test the hypothesis that dam removal would restore downstream hydrochory to levels observed in the upstream reach. To test this, we collected seeds in nets above and below the dam during three sample periods (early July, late July and early August), growing out seeds in a greenhouse and comparing seed abundance and species richness above and below dams, before and after dam removal. We found that after dam removal, the average number of hydrochorous seeds and species increased below Glines Canyon Dam to levels similar to or higher than that of the upstream reach; hydrochory levels in the upstream reach did not change. This study is the first to document the restoration of hydrochory in rivers following removal of a large dam. Restoration of hydrochory may ultimately increase downstream vegetation diversity and play a role in the recolonization of reservoir sediments deposited in the riparian zone in the years following dam removal. Copyright © 2016 John Wiley & Sons, Ltd.     

植被对河道水流及岸滩形态演变影响研究进展

植被作为河流系统重要的阻力因素,在河道形态演变方面发挥着至关重要的作用。植被地表结构改变了河道水流阻力、河床切应力、流速分布及水流紊动特性,植被根系增强了岸滩土体强度和稳定性。从植被地表茎干结构对水流特性影响和植被地下根系加筋作用对岸滩稳固影响两个方面的研究进行了回顾和综述,并指出今后在植被水流研究方面,可进一步考虑植被枝、叶对水流特性的影响,加强对柔性植被变形、摆动引起的动边界问题的研究,以及全面考虑河道形态边界条件的影响,特别是含植被的分汊河道水流特性还需深入研究;在植被根系固土护岸方面,可同时考虑岸滩土体非均质性、河道水流对岸滩的侵蚀冲刷作用及水体渗流对岸滩土体力学特性的影响;在植被河道形态演变预测方面,可以综合考虑含植被河道的水动力学及土力学问题,以及两者间的互馈响应机制。