Benthic macroinvertebrate assemblages in large river secondary channels: Contemporaneous and legacy effects of flow connectivity

In large rivers, secondary channels occur where the main channel is divided by an instream island, forming one or multiple smaller channels outside the main channel. Secondary channels are highly variable in morphometry, flow characteristics, and degree of connectivity to the main channel. Engineered closing structures at the upstream end of most secondary channels restrict inflow from the main channel, resulting in gradients of flow connectivity among secondary channels that vary with river stage. We investigated the relationship of flow connectivity to benthic macroinvertebrate assemblage richness and structure among a series of secondary channels of the Lower Mississippi River. Samples were collected over 2 years at times of high and low river stages. We discovered (1) macroinvertebrate assemblage structure and taxonomic richness varied along the flow connectivity gradient, and (2) there was a legacy effect of prior connection on assemblage structure that lasted up to a year. We contend that for management and restoration planning aimed at conservation of large river biological diversity, an important consideration are the life history requirements of animals utilizing secondary channel habitats.     

THE IMPORTANCE OF VARIABLE LATERAL CONNECTIVITY BETWEEN ARTIFICIAL FLOODPLAIN WATERBODIES AND RIVER CHANNELS

The rehabilitation of lowland rivers subjected to channelization and artificial levee construction should attempt to improve habitat heterogeneity and diversity of floodplain hydrological connectivity. However, rehabilitation efforts rarely consider the importance of variable lateral hydrological connectivity between floodplain waterbodies and main river channels (ranging from those permanently connected to those temporarily connected during river level rises), instead focusing on increasing individual floodplain waterbody connectivity. This study investigated the young‐of‐the‐year (YoY) fish communities in 10 artificial floodplain waterbodies of variable hydrological connectivity with the river Trent, England, between May and November 2006, inclusive. Floodplain waterbody connectivity to the main river was positively correlated with the number of species captured (alpha diversity), Shannon–Wiener diversity, Margalef's species richness index and the relative abundance of rheophilic species and negatively correlated with species turnover (beta diversity). YoY fish communities in poorly connected water bodies were most dissimilar to riverine communities. The results demonstrate the importance of variable lateral connectivity between artificial floodplain waterbodies and main river channels when rehabilitating lowland river fish communities. Copyright © 2011 John Wiley & Sons, Ltd.     

Effects of habitat restoration on the macroinvertebrate fauna in a foreland along the river waal,the main distributary in the Rhine delta

River engineering in the Rhine delta and water pollution have been major threats for the ecological functioning of the river in The Netherlands. To mitigate effects of river engineering, secondary channel construction in the forelands along the existing distributaries is considered to be an important measure for river restoration. These areas are the remnants of the former Rhine floodplain and the only area where habitat restoration is possible due to the river functions assigned. Secondary channel construction in the area called ‘Gamerensche Waarden’ was taken as an example to show effects of habitat restoration on the macroinvertebrate fauna. Totally 322 macroinvertebrate taxa were found during the monitoring period. During the first 3 years species richness in the area increased rapidly due to colonization processes in the channels following habitat development. After that period total number of taxa found in the channels stabilized at around 170. A clear positive relationship was demonstrated between habitat quality and species richness. Furthermore, the density of exotic species in the secondary channels was less than in the groyne fields of the main channel. The relatively low number of taxa in polluted habitats could be explained by the presence of the PCB 28 congener. Copyright © 2007 John Wiley & Sons, Ltd.     

Ecological connectivity in alluvial river ecosystems and its disruption by flow regulation

The dynamic nature of alluvial floodplain rivers is a function of flow and sediment regimes interacting with the physiographic features and vegetation cover of the landscape. During seasonal inundation, the flood pulse forms a ‘moving littoral’ that traverses the plain, increasing productivity and enhancing connectivity. The range of spatio-temporal connectivity between different biotopes, coupled with variable levels of natural disturbance, determine successional patterns and habitat heterogeneity that are responsible for maintaining the ecological integrity of floodplain river systems. Flow regulation by dams, often compounded by other modifications such as levee construction, normally results in reduced connectivity and altered successional trajectories in downstream reaches. Flood peaks are typically reduced by river regulation, which reduces the frequency and extent of floodplain inundation. A reduction in channel-forming flows reduces channel migration, an important phenomenon in maintaining high levels of habitat diversity across floodplains. The seasonal timing of floods may be shifted by flow regulation, with major ramifications for aquatic and terrestrial biota. Truncation of sediment transport may result in channel degradation for many kilometres downstream from a dam. Deepening of the channel lowers the water-table, which affects riparian vegetation dynamics and reduces the effective base level of tributaries, which results in rejuvenation and erosion. Ecological integrity in floodplain rivers is based in part on a diversity of water bodies with differing degrees of connectivity with the main river channel. Collectively, these water bodies occupy a wide range of successional stages, thereby forming a mosaic of habitat patches across the floodplain, This diversity is maintained by a balance between the trend toward terrestrialization and flow disturbances that renew connectivity and reset successional sequences. To counter the influence of river regulation, restoration efforts should focus on reestablishing dynamic connectivity between the channel and floodplain water bodies.     

Hydrologic connectivity and land cover affect floodplain lake water quality,fish abundance,and fish diversity in floodplain lakes of the Wabash-White River basin

Floodplain lakes are important aquatic resources for supporting ecosystem services, such as organismal habitat, biodiversity, and the retention of nutrients and sediment. Due to geomorphic alteration of river channels and land-cover change, degradation to floodplain lakes in the Ohio River basin is occurring at a rate that will escalate as climate change causes increased flood intensity and the seasonal redistribution of rainfall. A better understanding of the local drivers that affect oxbow lakes is needed for targeted floodplain restoration efforts designed to slow degradation. We examined the effects of land cover, topography, and hydrologic connectivity on water quality and fish diversity and abundance in nine floodplain lakes with potentially high remnant ecological function in the Wabash-White watershed (Indiana, Ohio, and Illinois). Data collection included water-quality parameters; stable water isotopes; total phosphorus, total nitrogen, and chlorophyll-a; and fish community diversity and abundance. Results indicate that hay/pasture land cover and decreased topographic relief in the local oxbow watersheds, along with reduced river hydrologic connectivity, were related to an increase in total phosphorus, total nitrogen, and chlorophyll-a. Greater biodiversity and abundance in fish assemblages were evident in oxbow lakes that were more disconnected from the main channel. The results of this study suggest that hydrologic connectivity of oxbow lakes with the contributing drainage area and the main channel influence nutrients and fish communities. Knowing the influencing factors can help ecosystem managers better protect these valuable floodplain lake ecosystems and prioritize restoration efforts amidst increasing stressors due to climate and land-use changes.     

HYDROLOGICAL CONNECTIVITY OF ABANDONED CHANNEL WATER BODIES ON A COASTAL PLAIN RIVER

Oxbow lakes, sloughs and other floodplain depressions associated with former channel positions are critical elements of floodplain hydrology, geomorphology and ecology. They comprise key elements of wetland and aquatic habitats and have important influence on the storage and routing of floodwaters. The hydrological connectivity between active river channels and floodplain depressions varies considerably in a qualitative sense, even within a single fluvial system. Several oxbows, sloughs and paleochannels were examined on the lower Sabine River, Texas/Louisiana, during a period of high but sub‐bankfull flow as well as at lower flows. Six different types of surface water connectivity with the main, active channel were identified: (i) flow through—a portion of the river flow regularly passes through the feature and returns to the main channel; (ii) flood channel—there is no hydraulic connection at normal flows, but at high flows the channels convey discharge, at least part of which returns to the main channel; (iii) fill and spill—the features fill to a threshold level at high flows and then overflow (mainly via ephemeral channels) into flood basins; (iv) fill and drain—the features fill at high river discharges but do not (except in large floods) overflow because as river discharge declines, water drains back to the river; (v) tributary occupied—tributaries draining to the abandoned channel continue to occupy it, flowing through it to the active channel; and (vi) disconnected—no flow is exchanged except during large floods. The age or stage of infilling and the relative elevation of abandoned channels are important first‐order controls of hydrological connectivity, but the lateral distance from the active channel is poorly related. Other critical controls are whether the cutoff section receives tributary input and whether a tie channel forms. The alluvial valley geomorphic context—specifically the presence of a meander belt ridge and flood basins—is also critical. Copyright © 2011 John Wiley & Sons, Ltd.     

Temporal changes in macroinvertebrate assemblages following experimental flooding in permanent and temporary wetlands in an Australian floodplain forest

The River Murray, Australia, is a highly regulated river from which almost 80% of mean annual flow is removed for human use, primarily irrigated agriculture. Consequent changes to the pattern and volume of river flow are reflected in floodplain hydrology and, therefore, the wetting/drying patterns of floodplain wetlands. To explore the significance of these changes, macroinvertebrate samples were compared between permanent and temporary wetlands following experimental flooding in a forested floodplain of the River Murray. Weekly samples from two permanent wetlands and four associated temporary sites were used to track changes in macroinvertebrate assemblage composition. Non‐metric multidimensional scaling was used to ordinate the macroinvertebrate data, indicating consistent differences between the biota of permanent and temporary wetlands and between the initial and later assemblages in the temporary sites. There were marked changes over time, but little sign that the permanent and temporary assemblages were becoming more alike over the 25‐week observation period. The apparent heterogeneity of these systems is of particular importance in developing river management plans which are likely to change flooding patterns. Such plans need to maintain a mosaic of wetland habitats if floodplain biodiversity is to be supported. Copyright © 2002 John Wiley & Sons, Ltd.     

Reconstruction of the characteristics of a natural alluvial river–floodplain system and hydromorphological changes following human modifications: the Danube River (1812–1991)

Based on detailed historical surveys from 1812, the natural riverine landscape of a 10.25‐km‐long reach of the Danube River in the Austrian Machland region prior to channelization is analysed. Anthropogenically induced changes of fluvial dynamics, hydrological connectivity and aquatic habitat composition are discussed, comparing the situations following channelization (1925) and flow regulation (1991). In 1812 the alluvial river–floodplain system of the Danube River comprised a highly complex channel network, numerous gravel bars and extensive islands, with the main channel and side arms (eupotamon) representing about 97% of the entire water surface at low flow. The floodplain was characterized by relatively flat terrain and numerous natural trenches (former active channels) connected to the main channel. These hydromorphological conditions led to marked expansion/contraction of the water surface area at water level fluctuations below bankfull (‘flow pulse’). The high degree of hydrological connectivity enabled intensive exchange processes and favoured migrations of aquatic organisms between the river and floodplain habitats over a period of approximately 90 days per year. Overall in 1812, 57% of the active zone (active channels and floodplain) was inundated at bankfull water level. Channelization and construction of hydropower plants resulted in a truncated fluvial system. Consequently, eupotamal water bodies decreased by 65%, and gravel/sand bars and vegetated islands decreased by 94% and 97%, respectively, whereas the area of the various backwaters doubled. In 1991 the former ‘flow pulse’ was halved due to artificial levees and embankments, greatly diminishing hydrological connectivity and decoupling large areas of the floodplain from the main channel. Active overflow, formerly playing an important role, is now replaced by backwater flooding and seepage inflow in isolated water bodies. Copyright © 2003 John Wiley & Sons, Ltd.     

A geomorphic monitoring and adaptive assessment framework to assess the effect of lowland floodplain river restoration on channel–floodplain sediment continuity

The state of the science of lowland river floodplain restoration reflects the relatively new and experimental nature of large river floodplain rehabilitation efforts. Based on results of a case study of floodplain restoration at the lowland Cosumnes River, California, we present a geomorphic monitoring and adaptive assessment framework that addresses the need to inform and utilize scientific knowledge in lowland floodplain river restoration activities. Highlighting hydrogeomorphic processes that lead to habitat creation, we identify a discharge threshold for connectivity and sediment transfer from the channel to the floodplain and integrate discharge magnitude and duration to quantify a threshold to aid determination of when geomorphic monitoring is warranted. Using floodplain sand deposition volume in splay complexes as one indicator of dynamic floodplain habitat, we develop a model to aid prediction of the sand deposition volume as an assessment tool to use to analyze future monitoring data. Because geomorphic processes that form the physical structure of a habitat are dynamic, and because the most successful restoration projects accommodate this fundamental characteristic of ecosystems, monitoring designs must both identify trends and be adapted iteratively so that relevant features continue to be measured. Thus, in this paper, adaptive assessment is defined as the modification of monitoring and analysis methods as a dynamic system evolves following restoration activities. The adaptive monitoring and assessment methods proposed facilitate long‐term measurements of channel–floodplain sediment transfer, and changes in sediment storage and morphology unique to lowland river–floodplain interactions and the habitat that these physical processes support. The adaptive assessment framework should be integrated with biological and chemical elements of an interdisciplinary ecosystem monitoring program to answer research hypotheses and to advance restoration science in lowland floodplain river systems. Copyright © 2005 John Wiley & Sons, Ltd.     

Short-term river response and restoration of biological diversity following slit construction

This study examined short-term temporal river restoration following slit construction using yearly surveys conducted from 2009 to 2011. The temporal changes caused by river restoration were monitored with regards to the river response, velocity diversity, channel geomorphic unit diversity and species diversity.The temporal change indicated a rapid increase in the hydraulic and channel geomorphic unit diversity by the river response, whereas the species diversity decreases by the rapid river response with the debris flow. The channel pattern changes were explained by an excess of shear stress, which eroded the bank toe. Bank scour or sediment failures then occurred during normal discharge. This process was the main mechanism of river widening in the Wasada stream.We suggested methods to assess the velocity and geomorphic diversity based on the Shannon diversity index for river conditions. The velocity and channel geomorphic unit diversity increased after the slit construction, with 1.31 in 2009, 1.68 in 2010, and 1.93 in 2011 for the velocity diversity and 1.05, 1.45, and 1.66 for the channel geomorphic unit diversity. Both diversities responded immediately to the slit construction, after which the response slowed. However, the species diversity remained lower than the pre-condition levels after the physical environment recovered. The reasons for the diversity decrease were the species evenness and the decrease in taxa richness. In the results, the species diversity varied as 2.33 (2009) to 2.38 (2010), and 2.12 (2011), while the species evenness decreased continuously: 0.79 (2009) to 0.74 (2010), and 0.73 (2011). The latter trend was caused by a rapid river response by debris flow that disturbed both the species population and species diversity. Species density and diversity decrease when the river response is very active in the early stage of river restoration.     

Temporal variations in organic carbon utilization by consumers in a lowland river

Understanding temporal trophic interactions in riverine food webs is essential for predicting river ecosystem function and improving management of these ecosystems. Temporal changes in energy flows through riverine food webs are readily assumed but are rarely tested. Temporal variability in food webs from two reaches of a lowland river (Ovens River, south‐eastern Australia) with differing levels of floodplain connectivity were examined over 12 months. We investigated how seasonal changes, flow variability and floodplain connectivity influence (i) stable isotope signatures of basal organic carbon sources (terrestrial sources: trees and grasses; aquatic sources: seston, biofilm and filamentous algae) and consumers (macroinvertebrates and fish) and (ii) the relative proportions of organic carbon sources contributing to consumer biomass using mixing models. We hypothesized (i) that during high flows, increased floodplain connectivity would increase the lateral exchange of terrestrial carbon subsidies to main channel consumers and (ii) that during low flow periods, main channel consumers would derive the majority of their carbon from aquatic benthic sources. Results indicated that isotope signatures for basal sources and for most of the consumers varied temporally and spatially. Mixing models indicated that increased floodplain connectivity did not increase terrestrial subsidies to consumers during high flows. Seston was the primary source during high flows whilst terrestrial vegetation increased in importance during low flows. Filamentous algae was also important during low flows for some consumers. These findings indicate that it is essential to include temporal variability in order to understand energy flows in lowland rivers, thus allowing for the dynamic nature of these ecosystems. Copyright © 2010 John Wiley & Sons, Ltd.     

Not all breaks are equal: Variable hydrologic and geomorphic responses to intentional levee breaches along the lower Cosumnes River,California

The transport of water and sediment from rivers to adjacent floodplains helps generate complex floodplain, wetland, and riparian ecosystems. However, riverside levees restrict lateral connectivity of water and sediment during flood pulses, making the re‐introduction of floodplain hydrogeomorphic processes through intentional levee breaching and removal an emerging floodplain restoration practice. Repeated topographic observations from levee breach sites along the lower Cosumnes River (USA) indicated that breach architecture influences floodplain and channel hydrogeomorphic processes. Where narrow breaches (<75 m) open onto graded floodplains, archetypal crevasse splays developed along a single dominant flowpath, with floodplain erosion in near‐bank areas and lobate splay deposition in distal floodplain regions. Narrow breaches opening into excavated floodplain channels promoted both transverse advection and turbulent diffusion of sediment into the floodplain channel, facilitating near‐bank deposition and potential breach closure. Wide breaches (>250 m) enabled multiple modes of water and sediment transport onto graded floodplains. Advective sediment transport along multiple flow paths generated overlapping crevasse splays, while turbulent diffusion promoted the formation of lateral levees through large wood and sediment accumulation in near‐bank areas. Channel incision (>2 m) upstream from a wide levee breach suggests that large flow diversions through such breaches can generate water surface drawdown during flooding, resulting in localized flow acceleration and upstream channel incision. Understanding variable hydrogeomorphic responses to levee breach architecture will help restoration managers design breaches that maximize desired floodplain topographic change while also minimizing potential undesirable consequences such as levee breach closure or channel incision.     

水系连通机理及其影响因素

为了摸清水系连通性衰退的成因,本文通过引入水系连通性指标和推求河道水力几何关系,深入分析了水系连通性的连通机理和影响因素.结果表明河道纵向连通性取决于河道来水来沙条件与边界形态,侧向连通性取决于河道分汇流比及分汇流区的水流流态和冲淤形态,河道横向连通包括河道滩槽并存、洪水期滩槽水沙交换以及水生植物的传播,主要是通过洪水...     

Aquatic macroinvertebrate assemblages of two contrasting floodplains: The rhǒne and ain rivers,France

The study examines the relationship between floodplain aquatic macroinvertebrates and sector-scale parameters such as geomorphology and history of regulation. The assemblages of six groups of invertebrates (Molluscs, Crustaceans, Ephemeroptera, Odonata, Trichoptera, and Coleoptera) were compared in various types of former channels from two contrasting but adjacent floodplain sectors: (1) the Jons sector of the Rhǒne River where successive meandering and braiding phases has left diversified fluvial forms in the landscape but where the main river is now embanked, and (2) the unregulated Ain River sector where the river is still actively meandering, although this process is being slowed by incision. The results demonstrate little difference in faunal composition between the two sectors but a significant difference in faunal structure. In the Rhǒne floodplain, there was a clear distinction between the faunal assemblages together with a high taxa richness at the scale of the sector (dominance of the beta diversity). In the Ain floodplain, the faunal assemblages were overlapping and the taxa richness was high at the sample scale (alpha diversity). Sector-scale spatial patterns of the faunal assemblages along former channels were also distinct: between-channel heterogeneity dominated in the Rhǒne, whereas within-channel heterogeneity dominated on the Ain. These results stress the influence of geomorphological and historical determinants on the floodplain communities and, conversely, the relevance of macroinvertebrate assemblages for the assessment, at the landscape scale, of aquatic systems within the floodplains.     

有植被的河道水流紊动特性模型试验研究

通过物理模型试验,研究了有植被的河道水流紊动特性.试验结果表明,在复式断面河道滩地种植柔性植被后,滩地糙率增大,水流紊动更为剧烈,河道水流紊动强度峰值由原先的滩槽交界区转移到滩地区.滩地的水流紊动强度沿程递减;滩槽交界区的水流紊动强度沿程不断增大;主槽的水流紊动强度主要与床面糙率有关,滩地植被影响了滩地水流的归槽时间,使主槽水流流速沿程增大.     

Effects of process-based floodplain restoration on aquatic macroinvertebrate production and community structure

Recent stream restoration approaches, such as Stage 0 restoration, aim to restore natural processes to regain lost ecosystem functions, but project implementation can also represent a reach-scale disturbance. Assumed outcomes of these restoration actions, like greater biological productivity, are rarely evaluated. In this study, we examined the short-term effects of Stage 0 floodplain restoration on the secondary production of aquatic macroinvertebrate communities in the South Fork McKenzie River, Oregon, 1–2 years following project implementation. We seasonally sampled macroinvertebrates from benthic and submerged wood surfaces in the restored reach, and two unrestored reference reaches located upstream, to estimate annual secondary production. Macroinvertebrate production estimates were 3× lower on a per-meter-squared basis in the restored reach than in the upstream unrestored reference reaches (9764 vs. 29,636 mg DM/m²/yr). However, because there was 4.5-times greater wetted area available in the restored reach, overall macroinvertebrate production per unit of valley length was 3.4× higher in the restored reach than in unrestored reference reaches (2744 vs. 802 kg DM/km). Additionally, the mosaic of aquatic habitats created via restoration (main-channel, side-channel, and wetted forest habitats) supported a diversity of macroinvertebrate assemblages both within and among reaches. Our findings suggest Stage 0 project implementation, which can include dewatering and filling incised channels, may reduce aquatic macroinvertebrate production on a per-unit-area basis for at least 1 or 2 years following restoration. However, this short-term disturbance effect may be offset by channel aggradation and widening, which can provide a more wetted area for macroinvertebrate production and may support greater macroinvertebrate community diversity. Future studies are needed to examine the longer term (2–10 years) aquatic macroinvertebrate response to Stage 0 restoration, and the impacts of shifting resource availability on stream fishes.     

Habitat loss as the main cause of the slow recovery of fish faunas of regulated large rivers in Europe: the transversal floodplain gradient

In large European rivers the chemical water quality has improved markedly in recent decades, yet the recovery of the fish fauna is not proceeding accordingly. Important causes are the loss of habitats in the main river channels and their floodplains, and the diminished hydrological connectivity between them. In this study we investigate how river regulation has affected fish community structure in floodplain waterbodies of the rivers Rhône (France), Danube (Austria), Rhine and Meuse (The Netherlands). A typology of natural and man‐made aquatic habitats was constructed based on geomorphology, inundation frequency and ecological connectivity, along the transversal river–floodplain gradient, i.e. perpendicular to the main stream of the river. Fish species were classified in ecological guilds based on their flow preference, reproduction ecology and diet, and their status on national red lists was used to analyse the present state of the guilds and habitats. Ecological fish guilds appear to be good indicators of ecological integrity and functioning of river–floodplain systems. A transversal successional gradient in fish community structure that bears some resemblance to the gradient found in natural rivers can still be discerned in heavily regulated rivers. It resembles the longitudinal river gradient; even some predictions of the River Continuum Concept are applicable. Overall, richness and diversity of species and ecological guilds decrease with decreasing hydrological connectivity of floodplain waterbodies. Anthropogenic disturbances have affected fish species unevenly: guilds of specialized species that are highly adapted to specifically riverine conditions have declined far more than generalist species. Fish habitats in the main and secondary channels have suffered most from regulation and contain the highest percentage of threatened species. Rheophilic fishes have become rare because their lotic reproductive habitats are severely degraded, fragmented, absent or unreachable. Limnophilic fishes have become rare too, mainly as a result of eutrophication. Eurytopic fishes have become dominant everywhere. Copyright © 2003 John Wiley & Sons, Ltd.     

Targeting lateral connectivity and morphodynamics in a large river‐floodplain system: The upper Rhine River

As a consequence of historical damming and channelization, most large rivers are disconnected from their floodplains, which therefore endure severe deficits in fluvial dynamics. Regaining some degree of lateral connectivity can lead to improved geomorphological and biological interactions. Yet, it is necessary to take into account limitations posed by current uses and legislation. This study presents a methodological approach to the selection of a realistic restoration target for a heavily modified large river segment, the free‐flowing Upper Rhine River downstream of Iffezheim dam (France–Germany border), based on the analysis of the existing biogeomorphic deficits, constraints set by human uses, and previous restoration experiences. To achieve the selected restoration target, proposed scenarios include embankment removal, bank lowerings, and side channel widenings with the aim of increasing lateral hydrological connectivity and promoting morphodynamics (bank erosion in lateral channels) that allow for the renewal of floodplain habitats. Results from 2‐D hydraulic simulations allow for a sensitivity analysis, comparing the current situation with the proposed scenarios, through parameters such as shore length of side channels actively connected at both ends to the main channel (eupotamon), and shear stress as a proxy for initiation of gravel erosion. Outcomes indicate that the two proposed restoration scenarios would succeed in reconnecting side channels and in increasing areas prone to substrate erosion, while maintaining flood protection and the heaviest navigation use among European rivers. The presented approach aids in the assessment of potential large river restoration scenarios and, thus, in the discussion of water management strategies.     

HYDROLOGIC CONNECTIVITY OF FLOODPLAINS,NORTHERN MISSOURI—IMPLICATIONS FOR MANAGEMENT AND RESTORATION OF FLOODPLAIN FOREST COMMUNITIES IN DISTURBED LANDSCAPES

Hydrologic connectivity between the channel and floodplain is thought to be a dominant factor determining floodplain processes and characteristics of floodplain forests. We explored the role of hydrologic connectivity in explaining floodplain forest community composition along streams in northern Missouri, USA. Hydrologic analyses at 20 streamgages (207–5827 km² area) document that magnitudes of 2‐year return floods increase systematically with increasing drainage area whereas the average annual number and durations of floodplain‐connecting events decrease. Flow durations above the active‐channel shelf vary little with increasing drainage area, indicating that the active‐channel shelf is in quasi‐equilibrium with prevailing conditions. The downstream decrease in connectivity is associated with downstream increase in channel incision. These relations at streamflow gaging stations are consistent with regional channel disturbance patterns: channel incision increases downstream, whereas upstream reaches have either not incised or adjusted to incision by forming new equilibrium floodplains. These results provide a framework to explain landscape‐scale variations in composition of floodplain forest communities in northern Missouri. Faust ( 2006 ) had tentatively explained increases of flood‐dependent tree species, and decreases of species diversity, with a downstream increase in flood magnitude and duration. Because frequency and duration of floodplain‐connecting events do not increase downstream, we hypothesize instead that increases in relative abundance of flood‐dependent trees at larger drainage area result from increasing size of disturbance patches. Bank‐overtopping floods at larger drainage area create large, open, depositional landforms that promoted the regeneration of shade‐intolerant species. Higher tree species diversity in floodplains with small drainage areas is associated with non‐incised floodplains that are frequently connected to their channels and therefore subject to greater effective hydrologic variability compared with downstream floodplains. Understanding the landscape‐scale geomorphic and hydrologic controls on floodplain connectivity provides a basis for more effective management and restoration of floodplain forest communities. Published 2013. This article is a U.S. Government work and is in the public domain in the USA.     

Fish assemblage patterns across a gradient of flow regulation in an Australian dryland river system

Hydrological regime, physical habitat structure and water chemistry are interacting drivers of fish assemblage structure in floodplain rivers throughout the world. In rivers with altered flow regimes, understanding fish assemblage responses to flow and physico‐chemical conditions is important in setting priorities for environmental flow allocations and other river management strategies. To this end we examined fish assemblage patterns across a simple gradient of flow regulation in the upper Murray–Darling Basin, Australia. We found clear separation of three fish assemblage groups that were spatially differentiated in November 2002, at the end of the winter dry season. Fish assemblage patterns were concordant with differences in water chemistry, but not with the geomorphological attributes of channel and floodplain waterholes. After the summer‐flow period, when all in‐channel river sites received flow, some floodplain sites were lost to drying and one increased in volume, fish assemblages were less clearly differentiated. The fish assemblages of river sites did not increase in richness or abundance in response to channel flow and the associated potential for increased fish recruitment and movement associated with flow connectivity. Instead, the more regulated river's fish assemblages appeared to be under stress, most likely from historical flow regulation. These findings have clear implications for the management of hydrological regimes and the provision of environmental flows in regulated rivers of the upper Murray–Darling Basin. Copyright © 2010 John Wiley & Sons, Ltd.