Effects of large woody debris on surface structure and aquatic habitat in forested streams,southern interior British Columbia,Canada

It is well known that large woody debris (LWD) plays an important functional role in aquatic organisms' life. However, the influence of LWD on channel morphology and aquatic environments at watershed levels is still unclear. The relationships between wood and surface structure and aquatic habitat in 35 first through fifth order streams of southern interior British Columbia were investigated. Study streams in the channel networks of the study watersheds were classified into four size categories based on stream order and bankfull width: Stream size I: bankfull width was less than 3 m, Stream size II: 3–5 m, Stream size III: 5–7 m, Stream size IV: larger than 7 m. We found the number of functional pieces increased with stream size and wood surface area in stream sizes I, II and III (24, 28 and 25 m²/100 m², respectively) was significantly higher than that in stream size IV (12 m²/100 m²). The contribution of wood pieces to pool formation was 75% and 85% in stream sizes II and III, respectively, which was significantly higher than those in stream size I (50%) and size IV (25%). Between 21% and 25% of wood pieces were associated with storing sediment, and between 20% and 29% of pieces were involved in channel bank stability in all study streams. Due to long‐term interactions, LWD in the intermediate sized streams (Size II and III) exhibited much effect on channel surface structure and aquatic habitats in the studied watersheds. Copyright © 2008 John Wiley & Sons, Ltd.     

Abundance and function of large woody debris in small,headwater streams in the Rocky Mountain foothills of Alberta,Canada

Large woody debris (LWD) significantly influences the structure and function of small headwater streams. However, what it contributes to geomorphic function depends on where it is located relative to the stream channel. We quantified LWD abundance and tested for associations among decay, position, orientation and function classes in 21 streams near Hinton, Alberta, Canada. LWD was more frequent (64.0 ± 3.3 LWD 100 m^?1) in streams in the Alberta foothills than it was in small streams in mountain, coastal, broadleaf deciduous and boreal forests, likely due to the narrow channel widths and low capacity of our study streams to transport logs downstream. LWD volumes were greater in coastal streams than in the Alberta foothills, likely due to differing tree sizes and decay rates. LWD morphology changed significantly as logs decayed and transitioned to different position and orientation classes. LWD in decay classes I and II were longest, most commonly in the bridge and partial bridge position classes, oriented perpendicular to stream flow, suspended above the channel and contributing least to stream geomorphic functions. LWD length and volume (but not diameter) decreased as decay advanced, making logs less stable. LWD in decay classes III and IV were strongly associated with partially bridged, loose, and buried position classes. They were more commonly diagonal or parallel to stream flow and contributed to bank stability, sediment retention, debris jams and riffle and pool formations. These results have been integrated into a conceptual model of LWD dynamics that provides a framework for future research on the mechanisms and rates of LWD recruitment, decay, transport and function. Copyright © 2010 John Wiley & Sons, Ltd.     

Experimental reintroduction of woody debris on the Williams River,NSW: geomorphic and ecological responses

A total of 436 logs were used to create 20 engineered log jams (ELJs) in a 1.1 km reach of the Williams River, NSW, Australia, a gravel‐bed river that has been desnagged and had most of its riparian vegetation removed over the last 200 years. The experiment was designed to test the effectiveness of reintroducing woody debris (WD) as a means of improving channel stability and recreating habitat diversity. The study assessed geomorphic and ecological responses to introducing woody habitat by comparing paired test and control reaches. Channel characteristics (e.g. bedforms, bars, texture) within test and control reaches were assessed before and after wood placement to quantify the morphological variability induced by the ELJs in the test reach. Since construction in September 2000, the ELJs have been subjected to five overtopping flows, three of which were larger than the mean annual flood. A high‐resolution three‐dimensional survey of both reaches was completed after major bed‐mobilizing flows. Cumulative changes induced by consecutive floods were also assessed. After 12 months, the major geomorphologic changes in the test reach included an increase in pool and riffle area and pool depth; the addition of a pool–riffle sequence; an increase by 0.5–1 m in pool–riffle amplitude; a net gain of 40 m³ of sediment storage per 1000 m² of channel area (while the control reach experienced a net loss of 15 m³/1000 m² over the same period); and a substantial increase in the spatial complexity of bed‐material distribution. Fish assemblages in the test reach showed an increase in species richness and abundance, and reduced temporal variability compared to the reference reach, suggesting that the changes in physical habitat were beneficial to fish at the reach scale. Copyright © 2004 John Wiley & Sons, Ltd.     

Invertebrate communities in Compensation Creek,a man‐made stream in boreal Newfoundland: The influence of large woody debris

Benthic invertebrate communities were examined in Compensation Creek, a man‐made stream in south‐central Newfoundland, Canada. Samples taken in September 2006 and September 2007 from large woody debris (LWD) were compared with samples from benthic environments to determine whether LWD supported a more diverse and abundant invertebrate community. Benthic habitats in a nearby natural stream were also sampled. Taxa composition was similar between the man‐made and the natural stream, highlighting successful colonization for the majority of taxa. Within Compensation Creek, taxa richness was higher in benthic habitats than on LWD, likely influenced by the successional age of the stream and surrounding habitat. The more complex benthic substrate provided refugia and allowed for the accumulation of fine detritus as a food source. Scrapers were almost completely absent from LWD and collector‐gatherer abundance was greater in the benthos. Collector‐filterer abundance was more than six times greater near the pond outflow than farther downstream when discharge was high, but abundances were almost equal when discharge was reduced. Riparian vegetation has not fully established around the man‐made stream, whereas it is overhanging and extensive at the natural stream, leading to more leaf‐litter input for shredders. As the morphology of Compensation Creek changes, the invertebrate community will continue to develop and likely increase utilization of accumulated detritus at LWD. Copyright © 2009 John Wiley & Sons, Ltd.     

A new methodology for the assessment of large woody debris accumulations on highly modified rivers (example of two French Piedmont rivers)

Methodologies that have been developed to quantify large woody debris (LWD) have been largely tested and adapted for mountain streams of the Pacific Northwest, characterised by a very high density of LWD, composed of large pieces of wood. In French rivers, LWD studies have focused on larger systems presenting low density and discrete distributions of LWD accumulations, where existing methods could not readily be used. We thus propose an easy-to-use method to quantify LWD within such systems. After defining three representative types of LWD, the volume is obtained by representing each LWD accumulation by a simple geometric form in order to measure its height, width and length. A model is then built for the different accumulation types to estimate wood mass from the measured volume. Since the measured volume is a combination of air and wood, we quantified the proportion of air, which is, respectively, equal to 18, 90 and 93% for trunks, wood jams and shrubs. To understand variability in wood mass, we evaluated the influence of different factors on wood density (defined as the ratio between mass and volume). The main factor was found to be the water absorption capacity of the wood, whereas a lesser factor was the degree of wood decay. Most wood pieces were found to increase their mass by an average of 100% and more after only 24 h in contact with water. Moreover, the observed levels of water loss and water absorption during the first 24 h of removal or exposure to water imply major short-term variations in wood mass, which may have significant consequences for wood transport during flooding. © 1998 John Wiley & Sons, Ltd.     

The role of riparian vegetation and woody debris in the development of macroinvertebrate assemblages in streams

Riparian vegetation development and macroinvertebrate assemblages were studied in 16 streams formed between 35 and 230 years ago, following glacial recession in Glacier Bay National Park, southeast Alaska. Riparian vegetation established most rapidly in streams where flow variation in downstream reaches was buffered by a lake. Riparian vegetation development was positively correlated with lower bank stability, but was independent of stream age. Roots and branches of riparian vegetation trailing into streams (trailing riparian habitat—TRH) were shown to be an important habitat for a number of macroinvertebrate taxa. In young and unstable streams, TRH was colonized mainly by Plecoptera whereas in more stable lake‐influenced streams Simuliidae dominated. Significant coarse woody debris (CWD) accumulations were not observed until after approximately 130 years of stream development had occurred when certain channel features, such as gravel bars, were stabilized by dead wood. Where dead wood was present, opportunistic wood taxa were abundant, even in the younger streams. However, a xylophagous species, Polypedilum fallax, was not recorded until streams were over 100 years old. Two‐way indicator species analysis (TWINSPAN) using presence/absence of macroinvertebrate taxa on TRH, initially divided streams into lake and non‐lake systems, but subsequent divisions were consistent with differences in stream age. TWINSPAN of macroinvertebrate assemblages on dead wood again highlighted differences in stream age. Canonical correspondence analysis indicated that bed stability and stream age were the most important environmental variables influencing macroinvertebrate distribution on TRH. Trailing riparian habitat was most abundant in moderately unstable streams where it facilitates invertebrate colonization. CWD contributes markedly to channel stabilization, provides habitat for invertebrate xylophages, and confers additional habitat complexity. Maximum levels of CWD are predicted to occur in non‐lake streams after approximately 300 years, but at least a further 100 years will be required in stable streams below lakes where dead wood entrainment is not enhanced by flooding, channel migration and bank undercutting. A conceptual model summarizing the role of TRH and CWD on stream development in Glacier Bay is presented. Copyright © 2005 John Wiley & Sons, Ltd.     

Large wood loads in an urban stream: The role of recruitment limitation versus transport dominance

Large wood (LW) has important physical and ecological functions in streams. Riparian vegetation is extensively removed during urban expansion, and urban streams may experience enhanced fluvial transport of LW due to flashy hydrology. In this study, LW loads were assessed for three reaches on North Buffalo Creek, an urban stream located in Greensboro, North Carolina, United States. These three reaches have similar hydrology but different riparian vegetation densities. We measured the frequencies and sizes of both in-channel LW and riparian vegetation across the three reaches. Our results showed that the recently reforested reach had greater LW volume (22.5 m³/km) compared to the unmanaged forested site (16 m³/km) and the site with low riparian vegetation density (4.78 m³/km). The difference in LW frequency among reaches was statistically significant ($p=.05$). However, the difference in the volume of individual pieces was not significantly different across reaches ($p=.84)$, indicating that a similar size of wood is recruited across the three sites. Our findings also showed that there is a positive relationship between riparian vegetation frequency and in-channel LW frequency, which are significantly related as a power function. Spatial lag models (integrating upstream riparian trees) did not show better results compared to a non-lagged model, suggesting that storage and recruitment were predominantly local and that the LW distribution at our reaches is limited by recruitment rather than dominated by fluvial transport. Our findings suggested that a fully forested watershed is not needed to provide some of the benefits of wood to urban streams.     

Macroinvertebrate community structure and function associated with large wood in low gradient streams

Large woody debris (wood) plays a number of important roles in forested stream ecosystems. Wood in streams provides habitat and flow refugia for fish and invertebrates, and is a site of biofilm production that serves as food for grazing organisms. Logs added to streams are rapidly colonized by invertebrates, and this habitat alteration is accompanied by changes in community composition and functional processes. A multiple habitat, qualitative sampling approach was employed to evaluate macroinvertebrate communities associated with wood habitats in 71 stream reaches in central Michigan and southeastern Minnesota, two agricultural regions in the midwestern United States. Macroinvertebrate taxa were classified with respect to behaviour (e.g. sprawler, clinger, swimmer), as well as trophic/feeding characteristics. These traits were used to examine community structure as a function of wood abundance and distribution. Although wood is not abundant in these streams and logs are generally small in size, wood is a very important habitat in both Michigan and Minnesota: 86% and 95% of the total taxa encountered at Michigan and Minnesota study sites, respectively, were found in wood habitats. Differences in regional patterns in the distribution of taxa across habitats were observed between Michigan and Minnesota. These are believed to result from differences in the number of habitat types available, and the dominant substrate composition. Local invertebrate diversity increased in Michigan, but not Minnesota, with the presence of wood habitats in streams. The presence of wood at a site increased the average taxa richness by 15 and 10 taxa in Michigan and Minnesota, respectively. Macroinvertebrate behavioural attributes and functional traits associated with wood habitats suggest that community traits may vary due to both local difference in flow and the location of wood in the channel. Copyright © 2003 John Wiley & Sons, Ltd.     

Bank stabilization,riparian land use and the distribution of large woody debris in a regulated reach of the upper Missouri River,North Dakota,USA

Large woody debris (LWD) is an important component of ecosystem structure and function in large floodplain rivers. We examined associations between LWD distribution and riparian land use, bank stabilization (e.g. riprap revetment), local channel geomorphology, and distance downriver from the dam in the Garrison Reach, a regulated reach of the upper Missouri River in North Dakota, USA. We conducted a survey of shoreline‐associated LWD in the reach during typical summer flow conditions. Reach‐wide LWD density was 21.3 pieces km^?1 of shoreline, of which most pieces (39% ) were ‘beached’ between the waterline and the bankfull level, 31% of pieces had evidence of originating at their current location (anchored), 18% of pieces were in deep water (>1 m), and 13% were in shallow water. LWD density along unstabilized alluvial (sand/silt) shorelines (27.3 pieces km^?1) was much higher than along stabilized shorelines (7.2 pieces km^?1). LWD density along forested shorelines (40.1 pieces km^?1) was higher than along open (e.g. rangeland, crop land; 9.2 pieces km^?1) or developed (e.g. residential, industrial; 7.8 pieces km^?1) shorelines. LWD density was highest overall along unstabilized, forested shorelines (45 pieces km^?1) and lowest along open or developed shorelines stabilized with a blanket‐rock revetment (5.5 pieces km^?1). Bank stabilization nearly eliminated the positive effect of riparian forest on LWD density. A predicted longitudinal increase in LWD density with distance from the dam was detected only for deep LWD (including snags) along unstabilized alluvial shorelines. Partial resurvey in the summer following the initial survey revealed a reduction in total LWD density in the reach that we attribute to an increase in summer flow between years. Changes in riparian management and land use could slow the loss of LWD‐related ecosystem services. However, restoration of a natural LWD regime in the Missouri River would require naturalization of the hydrograph and modification of existing bank stabilization and channel engineering structures. Copyright © 2004 John Wiley & Sons, Ltd.     

The effect of land use on channel geometry and sediment distribution in gravel mantled bedrock streams,Illinois River watershed,Arkansas

Headwater stream morphology is a direct reflection of watershed characteristics and therefore can inform our understanding of anthropogenic influence on channel geometry and sediment dynamics. Little knowledge of the geomorphology of headwater streams in the Ozark Plateaus region of northwest Arkansas exists. The Illinois River watershed, in northwest Arkansas, is of critical interest within the region because of land use changes in the headwaters due to rapid population growth. A mixture of forest and agricultural (open pasture and poultry houses) land use dominates the watershed, but urban areas are rapidly expanding. These land use types: forest, agriculture and urban are an effective proxy for increasing anthropogenic disturbance. Analysis of longitudinal profile, cross‐section and sediment distribution in streams from each land use type shows a strong trend of increasing slope and channel cross‐sectional area with a greater degree of anthropogenic disturbance. Additionally, urban streams are characterized by the presence of exposed bedrock in the stream bed, while agricultural and forested streams are gravel mantled. These data have important implications for current and future stream management policies and practices regionally. Copyright © 2010 John Wiley & Sons, Ltd.     

Hydrogeomorphic and Biotic Drivers of Instream Wood Differ Across Sub‐basins of the Columbia River Basin,USA

Instream wood promotes habitat heterogeneity through its influence on flow hydraulics and channel geomorphology. Within the Columbia River Basin, USA, wood is vital for the creation and maintenance of habitat for threatened salmonids. However, our understanding of the relative roles of the climatic, geomorphic, and ecological processes that source wood to streams is limited, making it difficult to identify baseline predictions of instream wood and create targets for stream restoration. Here, we investigate how instream wood frequency and volume differ between seven sub‐basins of the interior Columbia River Basin and what processes shape these differences within these sub‐basins. We collected data on wood volume and frequency, discharge and stream power, and riparian and watershed forest structure for use in modelling wood volume and frequency. Using random forest models, we found that mean annual precipitation, riparian tree cover, and the individual watershed were the most important predictors of wood volume and frequency. Within sub‐basins, we used linear models, finding that some basins had unique predictors of wood. Discharge, watershed area, or precipitation often combined with forest cover, riparian conifer, and/or large tree cover in models of instream large wood volume and frequency. In many sub‐basins, models showed at least one hydrologic variable, indicative of transport competence and one ecological variable, indicative of the reach or upstream watershed's capability to grow measurable instream wood. We conclude that basin‐specific models yield important insights into the hydrologic and ecological processes that influence wood loads, creating tractable hypotheses for building predictive models of instream wood. Copyright © 2015 John Wiley & Sons, Ltd.     

The introduction of woody debris into a channelized stream: effect on trout populations and habitat

Large woody debris was explored as a method of restructuring channelized streams to improve salmonid habitat. Whole trees were inserted in sections along a 2 km reach of a channelized stream to determine if large woody debris: (1) increased the abundance and biomass of brown (Salmo trutta) and rainbow trout (Oncorhynchus mykiss); (2) had an effect on physical habitat features; and (3) provided trouts with additional habitat. Trout populations and stream morphology were monitored before and after the introduction of woody debris and compared to control sections lacking woody debris. Abundance and biomass of both brown and rainbow trout increased in the treatment section compared to the control. Maximum and standard deviation of fish total length increased in all sections during summer months. The number of individuals and the standard deviations of total lengths decreased in the control section in winter, but increased in the treatment section. Mean water velocities decreased and number and volume of pools increased in treatment sections. Brown and rainbow trouts sought woody debris structures for cover. We conclude that large woody debris can serve as a method of reconstructing channelized streams to improve salmonid habitat. Copyright © 2002 John Wiley & Sons, Ltd.     

Large wood debris recruitment on differing riparian landforms along a Gulf Coastal Plain (USA) stream: a comparison of large floods and average flows

In southeastern Coastal Plain streams, wood debris can be very abundant and is recruited from extensive forested floodplains. Despite importance of wood debris, there have been few opportunities to examine recruitment and redistribution of wood in an undisturbed setting, particularly in the southeastern Coastal Plain. Following extensive flooding in 1994, measurements of individual downed trees (species, dbh, orientation, distance from base‐flow channel and condition) were made across replicated riparian landforms in a Gulf Coastal Plain 5th‐order stream. Annually, the fate of these trees was determined and newly recruited trees were noted. More than 300 downed trees have been recorded. Recruitment varied across landforms with more constrained reaches having greatest mortality. Total tree mortality varied substantially across years. Generally, tree recruitment was greatest in years with substantial floods (1994 and 1998). For each riparian landform type, tree mortality was correlated with the maximum daily flow during the period preceding annual debris surveys. This relationship was particularly strong for sand ridges (r² = 0.942) and low terraces (r² = 0.915), but was significant for floodplains (r² = 0.413). Greatest rates of debris recruitment per maximum daily flow were observed for sand ridges followed by low terraces. Flood characteristics also influenced debris recruitment. The 1994 flood was caused by a tropical storm and resulted in a rapid rise in streamflow. Much of the debris recruited during this flood was from toppled trees and was oriented parallel to the stream channel. In contrast, the 1998 flood was preceded by a wetter than average winter with more gradually rising flows and there was no relationship between riparian landform and debris characteristics. These results indicate that wood recruitment dynamics in Coastal Plain streams are complex. Wood recruitment rates are controlled by cyclical variations in climate interacting with riparian geomorphology. Infrequent high flows appear critical in the maintenance of the instream debris pool. Copyright © 2007 John Wiley & Sons, Ltd.     

The development of hydraulic and geomorphic complexity in recently formed streams in Glacier Bay National Park,Alaska

Geomorphic and hydraulic complexity within five streams representing 200 years of stream development were examined in Glacier Bay National Park, Alaska. Channel geomorphic units (CGUs) were mapped using a hierarchical approach, which defined stream habitat according to morphological and hydraulic characteristics. Detailed hydraulic assessment within the geomorphic units allowed differences in hydraulic characteristics across the 200‐year chronosequence to be documented. Channel geomorphology and hydrology changed as stream age increased. Younger streams were dominated by fast flowing geomorphic units such as rapids and riffles with little hydraulic or landscape diversity. As stream age increased, slower flowing habitat units such as glides and pools became more dominant, resulting in increased geomorphic, hydraulic and landscape diversity. These results suggest that geomorphic and hydraulic complexity develop over time, creating habitat features likely to be favoured by instream biota, enhancing biodiversity and abundance. Copyright © 2009 John Wiley & Sons, Ltd.     

Post‐flooding distribution and characteristics of large woody debris piles along the semi‐arid Sabie River,South Africa

The formation of large woody debris (LWD) piles has a profound impact on channel patterns and riparian succession in temperate rivers. The opportunity to study LWD along the Sabie River, a river in the semi‐arid region of Kruger National Park, South Africa, arose in February 2000 after a significant flood (c. 100‐year return interval) removed a large proportion of the fully mature riparian forest and other plant communities. Much of the uprooted vegetation was deposited as LWD piles (woody vegetation accumulations deposited on the ground > 0.1 m³) throughout the riparian and upland zones. In this article we describe the spatial distribution patterns of LWD as related to geomorphic channel type and flood frequency zone, and assess pile composition characteristics six months after the flood. Within the areas surveyed there were 68 LWD piles per hectare, the median size of LWD piles was 4.6 m³ but pile sizes (by volume) varied widely. Pool/rapid geomorphic channel types had the highest density of LWD piles (79 ha^?1) and the largest piles (by volume) were in the bedrock anastomosing channels (mean = 124 m³). Piles were larger in the seasonal and ephemeral flood frequency zones (mean = 54 m³ and 55 m³) than piles in the active zone (c. 2 m³). The patterns of distribution and volume of LWD will affect the subsequent development of vegetation communities as debris piles form a mosaic of patches of surviving organisms and propagules that can strongly influence the initial trajectory of succession. The amount, distribution, and subsequent decomposition of LWD are different from that reported for temperate rivers, suggesting that the role of LWD may be different on non‐floodplain rivers such as the Sabie in semi‐arid South Africa. Copyright © 2004 John Wiley & Sons, Ltd.     

Impact of a 100‐year flood on vegetation,benthic invertebrates,riparian fauna and large woody debris standing stock in an alpine floodplain

This paper investigates the impact of a 100‐year flood in May 1999 on community composition and large woody debris standing stock in an alpine floodplain (Isar, Germany). Detailed pre‐flood data sampled from 1993 to 1998 are compared with the situation directly after the flood. In those parts of the Isar floodplain mainly covered with pioneer vegetation prior to flooding, the coverage of unvegetated gravel bars increased by 22% following the flood. However, the flood did not remove larger amounts of older successional vegetation stages (willow thickets, floodplain forest). No significant changes in the benthic invertebrate fauna were recorded. The lowest densities of riparian ground beetles (Carabidae) within the study were recorded one month after the flood. Two months later, the ground beetle densities increased to the highest values ever recorded, indicating the ground beetle's high potential for recolonization. These results highlight the degree of resilience of both the aquatic and the riparian invertebrate fauna. The flood also caused a significant increase in large woody debris standing stock; in one section the number of logs increased tenfold and the volume increased by a factor of 20, leading to the assumption that most woody debris in alluvial flood‐plains is provided by catastrophic events. Copyright © 2004 John Wiley & Sons, Ltd.     

Spatial effects of reservoirs on fish assemblages in Great Plains streams in Kansas,USA

Reservoirs are important components of modern aquatic ecosystems that have negative impacts on native aquatic biota both up‐ and downstream. We used a landscape‐scale geographic information system (GIS) approach to quantify the spatial effects of 19 large reservoirs on upstream prairie fish assemblages at 219 sites in Kansas, USA. We hypothesized that fish assemblage structure would vary with increasing distance from a reservoir and that the abundance of reservoir fishes in upstream reaches would decline with distance from a reservoir. Ordination of sample sites showed variation in fish assemblage structure occurred primarily across river basins and with stream size. Variance partitioning of a canonical ordination revealed that the pure effect of reservoir distance explained a small but significant (6%; F = 4.90, P = 0.002) amount of variability in fish assemblage structure in upstream reaches. Moreover, reservoir species catch per unit of effort (CPUE) significantly declined with distance from a reservoir, but only in fourth‐ and fifth‐ order streams (r² = 0.32, P < 0.001 and r² = 0.49, P < 0.001, respectively). Finally, a multivariate regression model including measures of stream size, catchment area, river basin, and reservoir distance successfully predicted CPUE of reservoir species at sites upstream of Kansas reservoirs (R² = 0.45, P < 0.001). Overall, we found significant upstream effects of reservoirs on Kansas stream fish assemblages, which over time has led to a general homogenization of fish assemblages because of species introductions and extirpations. However, characteristic reservoir species are present throughout these systems and the importance of spatial proximity to reservoirs is probably dependent on the availability of suitable habitat (e.g. deep pools) in these tributary streams. Copyright © 2005 John Wiley & Sons, Ltd.     

Estimation of geomorphically significant flows in alpine streams of the Rocky Mountains,Colorado (USA)

Streamflows recorded at 24 gauging stations in the Rocky Mountains of Colorado were analyzed to derive regional regression equations for estimating the natural flow duration and flood frequency in reaches where the natural flows are unknown or have been altered by diversion or regulation. The principal objective of this analysis is to determine whether the relatively high, infrequent, but geomorphically and ecologically important flows in the Rocky Mountains can be accurately estimated by regional flow duration equations. The region considered in this study is an area of relatively abundant runoff, and, consequently, intense water resources development. The specific streams analyzed here, however, are unaltered and remain nearly pristine. Regional flow duration equations are derived for two situations. When the mean annual discharge is known, flows ≧10% of the time can be estimated with an uncertainty of ±9% for the 10% exceedance flow, to ±11% for the 1.0% exceedance flow. When the mean annual discharge is unknown, the relatively high, infrequent flow can be estimated using the mean basin precipitation rate (in m³/s), and basin relief with an uncertainty of ±23% for the 10% exceedance flow to ±21% for the 1.0% exeedance flow. The uncertainty in estimated discharges using the equations derived in this analysis is substantially smaller than has been previously reported, especially for the geomorphically significant flows which are relatively large and infrequent. The improvement is due primarily to the quality of streamflow records analyzed and a well‐defined hydrologic region.     

Hydraulic habitat composition and diversity in rural and urban stream reaches of the North Carolina Piedmont (USA)

The differences between urban and rural streams regarding hydrological process, channel morphodynamics and ecosystem functioning have been highlighted by a number of studies in recent decades. The need to understand lotic ecosystem functioning in these environments at scales relevant to individual organisms has led to research focusing on hydraulic composition and structure over small areas of channel bed. In this study we map and analyze the hydraulic biotope composition of two urban and two rural stream reaches in the North Carolina (USA) Piedmont to determine if urban flow regimes and attendant channel modification processes might translate into important differences in hydraulic environment, and if so, what those differences are. Hydraulic biotope assemblages were found to vary only moderately in diversity per unit stream length between sites, but were distinctly different in composition. One important control on the differences between rural and urban streams was found to be the localized incision of urban channels into bedrock and saprolite. Resistant rock outcrop in the beds of urban streams creates rapid and riffle biotopes and long stretches of upstream pool habitat by impoundment. Urban reaches were found to be more homogeneous than rural reaches in hydraulic composition and were dominated by pools. Rural reaches, characterized by copious sandy alluvium in the bed, were dominated by runs or glides. Quantitative differentiation of biotopes based on four hydraulic indices generally yields coherent associations, although these may vary in content. Comparisons between hydraulic and biotic diversities suggest relationships favoring biotic functional group aggregation over species richness‐based indices of diversity. Because the majority of published hydraulic diversity analyses are based on coarse‐bed streams, further study of hydraulic diversity in streams with finer substrate is likely to be beneficial. Copyright © 2008 John Wiley & Sons, Ltd.