Effect of aquatic mosses on juvenile fish density and habitat use in the regulated River Suldalslågen,western Norway

Two morphologically distinct moss communities were found in the River Suldalslågen. The liver moss community consists of species which form a dense mat on the bottom, while the river moss (Fontinalis) community forms long tufts. Moss growth has increased since hydropower regulations due to reduced floods and increased winter flows. Increased moss cover affects the bottom structure, as well as intra‐gravel and near‐bottom hydraulics. We studied densities of juvenile Atlantic salmon (Salmo salar) and brown trout (Salmo trutta) by electrofishing and habitat selection by direct underwater observation, in areas with natural moss cover compared with areas where mosses were experimentally removed. Areas with dense mats of liver mosses held lower densities of young of year (YoY) and older salmon parr than areas where liver moss had been removed. No differences in densities of YoY salmon were found between areas with and without Fontinalis. For older salmon, parr results were inconclusive. In some samples more and in others fewer fish were found in areas with Fontinalis moss removed. For trout, densities were higher in areas with Fontinalis, while results for liver moss were inconclusive. No major differences were found with regard to microhabitat selection between areas with and without river moss, suggesting that habitat quality in these areas was similar during summer, except with respect to substrate. Salmon held more exposed positions in areas without liver moss, but this is mainly attributed to different habitat availabilities. It is concluded that the relative increase in liver mosses in the River Suldalslågen has a negative impact on juvenile Atlantic salmon fish density. Copyright © 2002 John Wiley & Sons, Ltd.     

Contrasting temperatures,waterflows, and light: seasonal habitat selection by young Atlantic salmon and brown trout in a boreonemoral river

Habitat use and habitat selection by young Atlantic salmon and brown trout were investigated by direct underwater observation. We sampled during winter and summer water temperatures (low: 3–7°C; high: 9–12°C) coinciding with low and high waterflows (12–20 and 60–80 m³ s^?1), and during day and night in winter, and on six selected stations in the river. Observations of 396 salmon and 120 trout indicated a distinct seasonal pattern in behaviours and habitat selection. Feeding was the dominant behaviour at high water temperatures during summer. In winter, there was a diurnal pattern in behaviour; both species sheltered in interstitial spaces in the substrate during daylight, but during night held positions on or close to the substrate in slower flowing stream areas. Coarse substrate providing cover was therefore an important habitat factor during daylight at low water temperatures, while slow‐flowing water was important during night. Although spatial niche overlap was considerable both in summer and winter, salmon and trout segregated with respect to meso‐ and microhabitat selection, and relatively more at low temperatures. Both species changed their use of mesohabitats towards more slow‐flowing glide/flat habitats in winter. Irrespective of season, trout preferred in general more slow‐flowing water than salmon did, but the difference was more pronounced in winter. Salmon used a wider range of water depths and in particular water velocities, than did trout. Both species were less tolerant of high water velocities at low water temperatures. The seasonal and diurnal pattern in habitat selection reported have important implications for habitat research and habitat‐hydraulic modelling. Copyright © 2001 John Wiley & Sons, Ltd.     

Juvenile Salmonid Utilization of Floodplain Rearing Habitat After Gravel Augmentation in a Regulated River

Gravel augmentation is used in sediment‐starved streams to improve salmonid spawning habitat. As gravel is added to river channels, water surface elevations may rise in adjacent areas, activating floodplain habitat at lower flows, and floodplains inundate more frequently, potentially affecting the quantity and quality of juvenile salmonid rearing habitat. We analysed 5 years of juvenile Chinook salmon Oncorhynchus tschawytscha and steelhead Oncorhynchus mykiss data from snorkel surveys before and after gravel augmentation in the Lower American River, a low‐gradient, highly regulated alluvial river in California's Central Valley. We measured the quality and quantity of rearing habitat (current velocity and areal extent of inundated riparian vegetation) following gravel placement and tested whether these factors affected juvenile abundance. Gravel augmentation increased floodplain extent by 3.7–19.8%, decreased average flow velocity from 1.6 to 0.3 m s^?1 and increased the amount of vegetative cover from 0.3% to 22.6%. Juvenile abundances increased significantly for both species following augmentation. However, the strength of the relationship between abundance and habitat variables was greater for smaller salmonids. These results suggest that, in addition to enhancing salmonid spawning habitat, gravel augmentation can improve rearing habitat where channel incision and/or regulated hydrographs disconnect floodplains from main river channels. Copyright © 2015 John Wiley & Sons, Ltd.     

Population effect of a large‐scale stream restoration effort on Chinook salmon in the Pahsimeroi River,Idaho

Stream habitat restoration is an important tool for fisheries management in impaired lotic systems. Although small‐scale benefits of stream habitat restoration are commonly investigated, it is difficult to demonstrate population effects. The Pahsimeroi River Chinook salmon Oncorhynchus tshawytscha population was previously restricted to the lower portion of the river by multiple irrigation structures. To address fish passage issues, a combination of restoration projects was initiated including barrier removals, instream flow enhancements and installation of fish screens on diversions. The largest barrier was removed in 2009, more than doubling the amount of accessible linear habitat. We hypothesized restoration efforts would expand the distribution of spawning salmon in the Pahsimeroi River watershed, leading to a broader distribution of juveniles. We also hypothesized a broader juvenile distribution would have population effects by reducing the prevalence of density‐dependent growth and survival. Redds were documented in newly accessible habitat immediately following barrier removal and accounted for a median of 42% of all redds in the Pahsimeroi River watershed during 2009–2015. Snorkel surveys also documented juvenile rearing in newly accessible habitat. Juvenile productivity increased from a median of 64 smolts/female spawner for brood years 2002–2008 to 99 smolts/female spawner for brood years 2009–2014. Overall, results suggested increased habitat accessibility in the Pahsimeroi River broadened the distribution of spawning adult and rearing juvenile salmon and reduced the effects of density‐dependent survival. Large‐scale stream restoration efforts can have a population effect. Despite the large‐scale effort and response, habitat restoration alone is likely not sufficient to restore this population.     

Does ice matter? Site fidelity and movements by Atlantic salmon (Salmo salar L.) parr during winter in a substrate enhanced river reach

In‐stream habitat enhancement is a common remedial action in rivers where degradation/lack of suitable fish habitat can be diagnosed. However, post‐project monitoring to assess the response of the biota to modification is rare particularly during winter. We conducted in situ monitoring during the winters of 2004–2006 in the regulated Dalåa River, central Norway, in order to determine if winter habitat requirements of Atlantic salmon (Salmo salar L.) parr were realized in an enhanced (substrate and mesohabitat modification) reach. In total, 140 parr were marked with passive integrated transponder (PIT) tags and the fish were followed by carrying out active tracking surveys under variable ice conditions throughout the winter. Highest emigration (44%) occurred before ice formation started. Emigration was reduced after ice formed and was largely offset by parr re‐entering the enhanced area. Dispersal into the non‐enhanced, small substrate control area was observed only when the study reach was ice covered, and no parr were subsequently encountered in the control section after ice had melted. In the enhanced area, declining water temperature and surface ice conditions did not affect the spatial distribution of the resident salmon parr at the studied scale. Areas with ‘solid’ anchor ice precluded access for salmon parr whilst areas with ‘patchy’ anchor were used throughout the winter. Our results indicate that surface ice creates conditions that allow salmon parr to use stream habitats that otherwise provide only a limited amount of in‐stream cover. Ice processes should be taken into consideration when habitat enhancement projects are carried out and subsequently assessed for effectiveness. Copyright © 2008 John Wiley & Sons, Ltd.     

ESTIMATION OF JUVENILE SALMON HABITAT IN PACIFIC RIM RIVERS USING MULTISCALAR REMOTE SENSING AND GEOSPATIAL ANALYSIS

We conducted a regional classification and analysis of riverine floodplain physical features that represent key attributes of salmon rearing habitats. Riverine habitat classifications, including floodplain area and river channel complexity, were derived at moderate (30 m) spatial resolution using multispectral Landsat imagery and global terrain data (90 m) encompassing over 3 400 000 km² and most North Pacific Rim (NPR) salmon rivers. Similar classifications were derived using finer scale (i.e. ≤ 2.4‐m resolution) remote sensing data over a smaller set of 31 regionally representative flood plains. A suite of physical habitat metrics (e.g. channel sinuosity, nodes, floodplain width) were derived from each dataset and used to assess the congruence between similar habitat features at the different spatial scales and to evaluate the utility of moderate scale geospatial data for determining abundance of selected juvenile salmon habitats relative to fine scale remote sensing measurements. The resulting habitat metrics corresponded favorably (p < 0.0001) between the moderate scale and the fine scale floodplain classifications; a subset of these metrics (channel nodes and maximum floodplain width) also were strong indicators (R² > 0.5, p < 0.0001) of floodplain habitats defined from the finer scale analysis. These relationships were used to estimate the abundance and distribution of three critical shallow water floodplain habitats for juvenile salmon (parafluvial and orthofluvial springs, and shallow shore) across the entire NPR domain. The resulting database provides a potential tool to evaluate and prioritize salmon conservation efforts both within individual river systems and across major catchments on the basis of physical habitat distribution and abundance. Copyright © 2011 John Wiley & Sons, Ltd.     

The effect of wood and temperature on juvenile coho salmon winter movement,growth, density and survival in side‐channels

In British Columbia, side‐channels have been built to compensate for lost salmonid habitat. Most are structurally simple with little in‐stream wood; however, they support high densities of juvenile coho salmon. We longitudinally divided in halves the top 100 m of two dead‐end artificial side‐channels, one side‐channel with low winter water temperatures (surface‐fed) and one with relatively higher water temperatures (groundwater‐fed), closed the downstream end of each side‐channel with two‐way traps, and treated only one half of each channel with bundles of wood. Trapped fish were marked daily and coho salmon movement, growth and smolt output were monitored for two years. Wood addition increased juvenile coho winter carrying capacity and spring smolt output only in the ‘colder’ surface‐fed side‐channel. In contrast, in the groundwater‐fed side‐channel, with relatively higher water temperatures, the wood treatment slightly reduced the channel's carrying capacity and the spring output of coho salmon smolts. Copyright © 2003 John Wiley & Sons, Ltd.     

EVALUATING STATISTICAL APPROACHES TO QUANTIFYING JUVENILE CHINOOK SALMON HABITAT IN A REGULATED CALIFORNIA RIVER

Decisions on managed flow releases in regulated rivers should be informed by the best available science. To do this, resource managers require adequate information regarding the tradeoffs between alternative methodologies. In this study, we quantitatively compare two competing multivariate habitat models for juvenile Chinook salmon (Oncorhynchus tschawytscha), a highly valued fish species under serious decline in a large extent of its range. We conducted large‐scale snorkel surveys in the American River, California, to obtain a common dataset for model parameterization. We built one habitat model using Akaike Information Criterion analysis and model averaging, ‘model G’, and a second model by using a standard method of aggregating univariate habitat models, ‘model A’. We calculated Cohen's kappa, percent correctly classified, sensitivity, specificity and the area under a receiver operator characteristic to compare the ability of each model to predict juvenile salmon presence and absence. We compared the predicted useable habitat of each model at nine simulated river discharges where usable habitat is equal to the product of a spatial area and the probability of habitat occupancy at that location. Generally, model G maintained greater predictive accuracy with a difference within 10% across the diagnostic statistics. Two key distinctions between models were that model G predicted 17.2% less useable habitat across simulated flows and had 5% fewer false positive classifications than model A. In contrast, model A had a tendency to over predict habitat occupancy and under predict model uncertainty. The largest discrepancy between model predictions occurred at the lowest flows simulated and in the habitats most likely to be occupied by juvenile salmon. This study supports the utility and quantitative framework of Akaike Information Criterion analysis and model averaging in developing habitat models. Copyright © 2012 John Wiley & Sons, Ltd.     

Comparative analysis of glacial and nival streamflow regimes with implications for lotic habitat quantity and fish species richness

Growing interest in the differential responses of glacial and nival rivers to climatic forcing, and in ecological distinctions between the two streamflow regimes, suggests the need for a better comparative understanding of how the annual hydrologic cycle differs with presence or absence of catchment glacial cover. In this study, timing and magnitude characteristics of the average annual hydrographs of five glacierized and four nival catchments in the southwestern Canadian subarctic are empirically identified and compared. Likely effects upon fish habitat are qualitatively assessed, and net fisheries potential is tentatively investigated using taxa richness data. The chief hydrological conclusions at P < 0.05 using Kolmogorov–Smirnov and empirical orthogonal function analyses are: (1) catchment glacial cover results in freshets that are longer, larger, and peak later than those experienced by the nival regime; (2) the winter baseflows of glacial rivers are also much higher on a unit‐catchment‐area basis; and (3) basin scale and degree of catchment glacial cover are of comparable importance in determining the magnitude of the annual hydrologic cycle. These differences arise from the greater availability, both in volume and over time, of meltwater in glacial catchments, which in part reflects the consistently negative alpine glacial mass balances observed both in the present study area and globally under historical climatic warming. Such regime distinctions result in increased spawning season and winter aquatic habitat availability, which may in turn offset negative habitat characteristics previously identified for glacial river ecosystems. While previous studies have suggested that glacial influences tend to decrease macroinvertebrate diversity and increase salmon populations, preliminary analysis of available fish species presence/absence data from the current study area tentatively appears to suggest similar or, perhaps, slightly higher fish taxa richness relative to nival streams; in all three cases, however, catchment lake cover may play a key hydroecological modifying role. The results strongly confirm and extend existing understanding of glacial–nival regime differences with respect to both streamflow and fisheries ecology, and raise new questions for future research. Copyright © 2005 John Wiley & Sons, Ltd.     

Seasonal selection of habitat by Spotted Bass and Shorthead Redhorse in a regulated river in the Midwest,USA

Riverine fish populations depend on habitats supporting their resource and life history needs. Dynamic streamflow caused by river regulation or natural events influences the distribution of downstream habitat characteristics. Through studying habitat selection, we can identify the most utilized and valuable habitats for the success of native fishes. We determined seasonal habitat selection of two common, native fish species on the Osage River downstream of Bagnell Dam, a hydroelectric dam in central Missouri, from April 2016 to June 2017 using radio telemetry. Spotted Bass (Micropterus punctulatus) are nest‐guarders, sight feeders, and habitat generalists, whereas Shorthead Redhorse (Moxostoma macrolepidotum) are fluvial dependent, migratory, and benthic feeders. Bayesian discrete choice analyses determined that both species selected particular water depth, velocity, and presence of submerged cover in some or all seasons, even as available habitat changed. Spotted Bass selected water depths <4.0 m near submerged cover during all seasons, low velocity during spring and summer, and near‐bank habitat in all seasons except spring. Shorthead Redhorse used fast flowing habitat during spring, 0.4–1.1 m/s velocity during summer, and low velocity in fall and winter (0.1–0.5 m/s). Shorthead Redhorse used submerged cover in all seasons except summer and selected specific ranges of depth within spring (2.4–4.4 m), summer (3.3–6.7 m), and winter (1.1–2.3 m). Our findings suggest that maintaining habitats with cover and diverse water depths and velocities, particularly both low and high velocity habitats during spring, may promote resilience by providing beneficial habitats for native fishes.     

Fish assemblages and stream hydraulics: consistent relations across spatial scales and regions

General relationships between organisms and their habitat, consistent across spatial scales and regions, suggest the existence of repeatable ecological processes and are useful for the management of stream networks. From published data, we defined four guilds of European fish species with contrasting preferences for microhabitat hydraulics within stream reaches. At the scale of stream reaches and across 139 French sites (590 460 fishes sampled), we analysed how fish guild proportions were related to reach hydraulics (proportion of pools vs. riffles %POOL; median discharge by unit width Q50/W). The strongest correlations were observed between two fish guilds and %POOL (p < 0.001, r² ≥ 0.41) and between one fish guild proportion and Q50/W (p < 0.001, r² = 0.10). These reach–scale relationships were consistent across six large French basins, and consistent with the analyses made at the microhabitat scale. Therefore, microhabitat preferences for hydraulics are strong enough to generate consistent reach‐scale community responses to hydraulics across regions, despite the influence of other filters such as temperature, nutrient levels or history. The distribution of basic geomorphic features (pools, riffles) in streams and their modification (by dams, weirs and dikes) can modify the proportion of fish guilds by up to 80%, probably contributing to the long‐term decline of riffle‐dwelling species in Europe. Copyright © 2006 John Wiley & Sons, Ltd.     

Comparison of MesoHABSIM with two microhabitat models (PHABSIM and HARPHA)

This study demonstrates how using different habitat models can influence the results of instream habitat assessment and conclusions for river management. We used three models for a portion of the Quinebaug River (Connecticut and Massachusetts, USA): a simplistic microhabitat model with univariate habitat‐use criteria and substrate‐based channel index (the Physical Habitat Simulation Model (PHABSIM)); a microhabitat model using multivariate criteria including a wide range of cover attributes (HARPHA); and a mesohabitat model with multivariate habitat‐suitability criteria (MesoHABSIM). The flow‐habitat rating curves produced by each model were compared at two scales: site and study segment. To investigate the impact of model choice on answering questions such as which location or flow provides more habitat, we applied Spearman's correlation of ranks. The relationship between habitat‐suitability predictions and fish presence at the same location was investigated with dedicated fish observations. The study showed that: (1) of the tested models, only MesoHABSIM predictions correlated with fish observations; (2) the variation within microscale models (PHABSIM and HARPHA) was greater than between micro‐and mesoscale models (HARPHA and MesoHABSIM); and (3) simple univariate habitat‐use criteria provided the largest source of discrepancies among the models. We suggest that these differences may lead to erroneous conclusions, especially if flow‐habitat rating curve analysis is considered an endpoint of instream flow study. Copyright © 2007 John Wiley & Sons, Ltd.     

EVALUATION OF RIPARIAN HABITAT RESTORATION IN A LOWLAND RIVER

For many years, navigable lowland rivers have been embanked artificially or suffered from substantial shipping wave action, leading to habitat degradation. Recently, riparian habitats were restored by creating foreshores and spawning grounds in the river Yser, a lowland river in Flanders, Belgium. The aim of this paper was to evaluate the role of these restored habitats for spawning and nursery of juvenile fish. To cover a wide range of anthropogenic disruption, four riparian mesohabitat types were selected and compared, ranging from semi‐natural over artificial spawning grounds and foreshores to artificial embankments. Juvenile fish were subjected to sampling by using electrofishing between June and September 2009 at different microhabitats located in five sites of each riparian mesohabitat type. Juvenile fish strongly preferred natural riparian habitats, whereas artificial embankments showed the lowest species richness, abundance and functional organization of juvenile fish species. Restored riparian habitats appeared to be an appropriate alternative for artificial embankments in navigable lowland rivers but still score significantly less than natural habitats. Juvenile fish avoided bare microhabitats but did not prefer any other microhabitat type (reed, woody or grassy vegetation), emphasizing the importance of microhabitat diversity. This paper provides valuable insights into riparian habitat restoration to river managers and stakeholders. Copyright © 2011 John Wiley & Sons, Ltd.     

Effects of artificial woody structures on Atlantic salmon habitat and populations in a Nova Scotia stream

A 1‐km reach of Brierly Brook, Nova Scotia, was studied from 1995 to 2004 to determine if the addition of artificial structures mimicking large woody debris could enhance Atlantic salmon populations. In 1995, digger logs (which mimic fallen trees) and deflectors (which narrow the channel) were constructed in a 250‐m section of the brook devoid of woody debris (Old Restored Site). In 2003, 5 more digger logs and defectors were built in a previously unrestored section of the stream (New Restored Site). A third control site was left unchanged. Physical changes caused by the structures were monitored at the New Restored Site. Densities of juvenile and spawning Atlantic salmon were also monitored. At all sites, woody debris structures in the brook were important and effective in creating complex salmonid habitat. The structures narrowed the channel, scoured pools and undercut banks. They created habitat that parr used for summer and winter refuge and adult spawners used for cover and resting during upstream migration and spawning. The structures caused gravels to accumulate that spawning adults used to build redds and fry used for shelter. The reaches with structures had higher spawning densities than reaches without them; spawning increased in the New Restored Site relative to the control site. The absence of woody debris may be a bottleneck for salmonid populations in streams of the Atlantic Northeast. For streams with a small or immature riparian zone and little woody debris in the channel, woody structures may be an effective tool for restoring salmonid populations. Copyright © 2008 John Wiley & Sons, Ltd.     

Behavioural responses of juvenile Atlantic salmon (Salmo salar) to presence of boulders

The influence of boulder presence on the behaviour of juvenile Atlantic salmon (Salmo salar) was investigated experimentally in an indoor flume fed by a 16 l s^?1 flow of natural river water. The flume was divided into 16 arenas (each 1 m²) that were landscaped with river gravel and standardized boulders, to represent relative ‘complex’ or ‘simple’ habitats. Each arena housed three wild‐caught fish. In a three‐week trial, the effects of landscape on aspects of individual behaviour were recorded. Food intake was highest in the simple landscape and directly related to social status and time spent in the water column. The fish in the complex chambers actively maintained station in the water column significantly more than fish in simple landscapes and therefore partially compensated for reduced foraging rates associated with complexity. Fish in simple chambers spent more time orientated upstream than those in complex landscapes. There was no evidence that habitat complexity influenced levels of aggression, average aggressive distance, ‘constrained’ territory size, or dominance. Activity and space use varied with social status. Overall, this study illustrates that addition of boulders can result in costs to Atlantic salmon parr, which can be expected to offset to some extent benefits brought about by increased stream complexity. Work is now needed to evaluate the balance of costs and benefits at different life stages and under various environmental conditions. © Crown copyright 2005. Reproduced with the permission of Her Majesty's Stationery Office. Published by John Wiley & Sons, Ltd.     

Response of Fish Communities to Hydrological and Morphological Alterations in Hydropeaking Rivers of Austria

Climate change asks for the reduction in the consumption of fossil‐based fuels and an increased share of non‐regulated renewable energy sources, such as solar and wind power. In order to back up a larger share of these intermittent sources, ‘battery services’ are needed, currently provided only in large scale by hydropower, leading to more rapid and frequent changes in flows (hydropeaking) in the downstream rivers. Increased knowledge about the ecosystem response to such operations and design of cost‐effective measures is needed. We analysed the response of fish communities to hydropeaking (frequency, magnitude, ramping rate and timing) and the interaction with the habitat conditions in Austrian rivers. An index of biotic integrity (Fish Index Austria) was used to compare river sections with varying degrees of flow fluctuations under near‐natural and channelized habitat conditions. The results showed that habitat conditions, peak frequency (number of peaks per year), ramping rate (water level variation) and interaction between habitat and ramping rate explained most of the variation of the Fish Index Austria. In addition, peaking during the night seems to harm fish more than peaking during the day. Fish communities in hyporhithral and epipotamal types of rivers are more affected by hydropeaking than those in metarhithral type of rivers. The results support the findings of other studies that fish stranding caused by ramping rates >15 cm h^?1 are likely to be the main cause of fish community degradation when occurring more often than 20 times a year. While the ecological status degrades with increasing ramping rate in nature‐like rivers, fish communities are heavily degraded in channelized rivers regardless of the ramping rate. The mitigation of hydropeaking, therefore, requires an integrative approach considering the combined effects of hydrological and morphological alterations on fish. © 2014 The Authors. River Research and Applications published by John Wiley & Sons, Ltd.     

Juvenile coho salmon behavioural characteristics in Klamath river summer thermal refugia

During the summer in the main‐stem Klamath River, juvenile salmonids respond positively to cooler tributary temperatures by congregating in large schools at the mouths of these tributaries, referred to as thermal refugia. The purpose of this paper is to summarize results from coho salmon (Oncorhynchus kisutch) thermal refugia studies conducted since 2006 in the Klamath River. Results showed that juvenile coho salmon started using thermal refugia when the Klamath River main‐stem temperature approached approximately 19 °C. The majority of the juvenile coho salmon within the studied thermal refugia were found in the slower velocity habitat associated with cover. Juvenile coho salmon counts in the studied thermal refugia dramatically decreased at temperatures >22–23 °C, suggesting that this approximates their upper thermal tolerance level. Although some juvenile coho salmon were very mobile, others chose, until smoltification, to rear in the mouths and lower reaches of non‐natal tributaries where thermal refugia occurred, apparently because of suitable habitat conditions. Information gained from these investigations will improve our knowledge of the life history of coho salmon in the Klamath River drainage and how they use the main‐stem river. Copyright © 2010 John Wiley & Sons, Ltd.     

Seasonal Rearing Habitat in a Large Mediterranean‐Climate River: Management Implications at the Southern Extent of Pacific Salmon (Oncorhynchus spp.)

Pacific salmon (Oncorhynchus) use a variety of rearing environments prior to seaward migration, yet large river habitats and their use have not been well defined, particularly at the southernmost salmon range where major landscape‐level alterations have occurred. We explored juvenile Chinook salmon (Oncorhynchus tshawytscha) and steelhead (Oncorhynchus mykiss) presence along the river continuum and in main‐channel and off‐channel habitats of a regulated California Mediterranean‐climate river. Over an 8‐year period, off‐channels of the lower Mokelumne River exhibited slower and warmer water than the main‐channel. Probability of salmonid presence varied by stream reach and habitat types. Steelhead and Chinook salmon both demonstrated transitional responses to the dry season, with juveniles leaving off‐channels by midsummer. This corresponded to flow recession, increasing water temperatures, salmonid growth and end of emigration period. Main‐channel steelhead observations continued until the following storm season, which brought cool flood flows to reconnect off‐channels and the next juvenile cohort of both species to the river. Within arid climates, low‐gradient off‐channels appear more transiently used than in cooler and more northern humid climate systems. Within a highly regulated Mediterranean‐climate river, off‐channel habitats become increasingly scarce, disconnected or temperature limiting in low‐gradient reaches both seasonally and due to anthropogenic modifications. These observations may provide guidance for future management within large salmon streams. Copyright © 2015 John Wiley & Sons, Ltd.     

Interactive effects of cover and hydraulics on brown trout habitat selection patterns

Habitat modelling results are extremely sensitive to the habitat suitability criteria (HSC) used in the simulations. HSCs are usually expressed as univariate habitat suitability curves, although such univariate approach has been long questioned, since overlooking interactions between hydraulic variables may misrepresent the complexity of fish behaviour in habitat selection. It could lead to adopt erroneous flow management decisions based on misleading results. Furthermore, the interactive effects of hydraulic variables on habitat selection may be driven by the structural features of the channel, which determine cover availability. Therefore, we compared brown trout habitat selection patterns through multivariate resource selection functions (RSFs) in structurally contrasting rivers to unveil the interactive effects of hydraulics and cover elements and their consequences in univariate HSC results. Microhabitat preferences of young‐of‐the‐year (0+) trout were similar across fast and slow waters, meanwhile juvenile (1+) and adult (>1+) preferences significantly changed. RSFs for young‐of‐the‐year trout were consistent with univariate results and did not differ among water types. However, RSFs for older trout varied among water types and revealed complex interactions among hydraulic variables and between hydraulics and structural elements, which were not described accurately by univariate curves. Therefore, results suggest that interactions between water depth and current velocity have a significant effect on habitat selection patterns in juvenile and adult brown trout, this effect being controlled by cover availability. Copyright © 2008 John Wiley & Sons, Ltd.