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1.
The performance and movements of juvenile Atlantic salmon Salmo salar exposed to variable water discharge (simulating hydropeaking) but with a stable water‐covered area were studied in six experimental stream channels, both during the winter and summer. Thirty fish were stocked into each channel, and the growth, body fat and movements of the fish were followed for about 2.5 months in each season. During the winter, no effect of hydropeaking was documented on performance or movement. In the summer, fish experiencing hydropeaking had lower body mass, lower body fat, and higher movement rates than the control fish. In general, effect sizes were small, and the rapid and frequent changes in water discharge and water level in the present study had small effects on the performance of juvenile Atlantic salmon. The cumulative long‐term effect at the population level is unknown, but a reduced growth rate of 10% and a reduction in body fat of 16% in the hydropeaking experiments in the summer might to some extent translate into increased smolt age and lower overwintering survival. Copyright © 2014 John Wiley & Sons, Ltd.  相似文献   

2.
Elevated stream temperature is a primary factor limiting the coho salmon (Oncorhynchus kisutch) population in California's Shasta River Basin. Understanding the mechanisms driving spatial and temporal trends in water temperature throughout the Shasta River is critical to prioritising river restoration efforts aimed at protecting this threatened species. During the summer, the majority of streamflow in the Shasta River comes from large‐volume, cold‐water springs at the head of the tributary Big Springs Creek. In this study, we evaluated the initial character of this spring water, as well as the downstream fate and transport of these groundwater inflows during July and August 2008. Our results indicated that Big Springs Creek paradoxically provided both cool and warm waters to the Shasta River. During this period, cool groundwater inflows heated rapidly in the downstream direction in response to thermal loads from incoming solar radiation. During the night time, groundwater inflows did not appreciably heat in transit through Big Springs Creek. These diurnally varying water temperature conditions were inherited by the Shasta River, producing longitudinal temperature patterns that were out of phase with ambient meteorological conditions up to 23 km downstream. Findings from this study suggest that large, constant temperature spring sources and spring‐fed rivers impart unique stream temperature patterns on downstream river reaches that can determine reach‐scale habitat suitability for cold‐water fishes such as coho salmon. Recognising and quantifying the spatiotemporal patterns of water temperature downstream from large spring inflows can help identify and prioritize river restoration actions in locations where temperature patterns will allow rearing of cold‐water fishes. Copyright © 2013 John Wiley & Sons, Ltd.  相似文献   

3.
Juvenile salmonids can become stranded over gravel bars or trapped in off-channel habitat during rapid flow decreases that often occur in regulated rivers. In a stream channel experiment that simulated stranding over a gravel bar, more juvenile chinook salmon (Oncorhynchus tshwytscha) were stranded when the water was 6°C compared with 12°C. The rate of flow decrease was not a significant factor in the incidence of stranding. In contrast, the number of chinook and coho (O. kisutch) salmon juveniles that became trapped in side channels built in the stream channel increased with increasing rate of dewatering, and for coho salmon, more fish were trapped at night compared with the day. Even at the slowest rate of flow decrease some fish remained in the side channels after the channels became disconnected from the main flow. My results suggest that mortality as a result of trapping in side channels or pot-holes will be decreased, but may not be eliminated, by flow ramping, although this result needs to be corroborated with field studies. © 1997 John Wiley & Sons, Ltd.  相似文献   

4.
Pacific salmon (Oncorhynchus) use a variety of rearing environments prior to seaward migration, yet large river habitats and their use have not been well defined, particularly at the southernmost salmon range where major landscape‐level alterations have occurred. We explored juvenile Chinook salmon (Oncorhynchus tshawytscha) and steelhead (Oncorhynchus mykiss) presence along the river continuum and in main‐channel and off‐channel habitats of a regulated California Mediterranean‐climate river. Over an 8‐year period, off‐channels of the lower Mokelumne River exhibited slower and warmer water than the main‐channel. Probability of salmonid presence varied by stream reach and habitat types. Steelhead and Chinook salmon both demonstrated transitional responses to the dry season, with juveniles leaving off‐channels by midsummer. This corresponded to flow recession, increasing water temperatures, salmonid growth and end of emigration period. Main‐channel steelhead observations continued until the following storm season, which brought cool flood flows to reconnect off‐channels and the next juvenile cohort of both species to the river. Within arid climates, low‐gradient off‐channels appear more transiently used than in cooler and more northern humid climate systems. Within a highly regulated Mediterranean‐climate river, off‐channel habitats become increasingly scarce, disconnected or temperature limiting in low‐gradient reaches both seasonally and due to anthropogenic modifications. These observations may provide guidance for future management within large salmon streams. Copyright © 2015 John Wiley & Sons, Ltd.  相似文献   

5.
During the summer in the main‐stem Klamath River, juvenile salmonids respond positively to cooler tributary temperatures by congregating in large schools at the mouths of these tributaries, referred to as thermal refugia. The purpose of this paper is to summarize results from coho salmon (Oncorhynchus kisutch) thermal refugia studies conducted since 2006 in the Klamath River. Results showed that juvenile coho salmon started using thermal refugia when the Klamath River main‐stem temperature approached approximately 19 °C. The majority of the juvenile coho salmon within the studied thermal refugia were found in the slower velocity habitat associated with cover. Juvenile coho salmon counts in the studied thermal refugia dramatically decreased at temperatures >22–23 °C, suggesting that this approximates their upper thermal tolerance level. Although some juvenile coho salmon were very mobile, others chose, until smoltification, to rear in the mouths and lower reaches of non‐natal tributaries where thermal refugia occurred, apparently because of suitable habitat conditions. Information gained from these investigations will improve our knowledge of the life history of coho salmon in the Klamath River drainage and how they use the main‐stem river. Copyright © 2010 John Wiley & Sons, Ltd.  相似文献   

6.
We examined the relationship between the physical environment and habitat use of juvenile masu salmon, Oncorhynchus masou, in the Nobori River in Hokkaido, Japan to provide a perspective for the conservation of fish habitat in regulated streams. The study was undertaken during the autumn and winter, with an emphasis on the hierarchy of three spatial scales: microhabitat, channel‐unit and reach scales. The microhabitat‐scale analysis indicated juvenile masu salmon preferred a midstream habitat type, with a greater depth (Avg. ± SD: 35.4 ± 14.2 cm) and high (43.4 ± 23.1 cm s?1) and uniform current velocities during the autumn, and a channel margin habitat type with a moderate current (about 20 cm s?1) and submerged cover during winter. In addition, different cover types have different roles in determining juvenile salmon distributions during winter. Grass cover had extremely high carrying capacities, whereas coarse substrate cover provided winter habitat for larger juvenile salmon. Channel‐unit scale analyses showed that abundance of juvenile salmon tended to be higher in pools than runs in the autumn through winter. Reach‐scale analysis showed that abundance and mean body length of juvenile salmon significantly differed between differently regulated reaches during winter, associated with the dominant cover type in each reach. This study demonstrated that the habitat conditions determining juvenile masu salmon distribution differ according to the season and scale of analysis. Therefore, for conservation of fish communities, it is important to evaluate and conserve or create fish habitats in regulated reaches, with a focus on the hierarchy of spatial scales and seasonal differences. Copyright © 2007 John Wiley & Sons, Ltd.  相似文献   

7.
Engineered log jams (ELJs) are increasingly being used in large rivers to create fish habitat and as an alternative to riprap for bank stabilization. However, there have been few studies that have systematically examined how juvenile salmonids utilized these structures relative to other available habitat. We examined Chinook salmon (Oncorhynchus tshawytscha), coho salmon (O. kisutch) and trout (O. mykiss and O. clarki) response to the placement of engineered log jams (ELJs) in the Elwha River, Washington State, USA. We used summer snorkel surveys and a paired control‐treatment design to determine how engineered log jams in a large river system affect the density of juvenile salmon. We hypothesized that densities of juvenile salmonids would be greater in habitats with ELJs than in habitats without ELJs in the Elwha River and that this ELJ effect would vary by species and size class. Juvenile salmonid density was higher in ELJ units for all control‐treatment pairs except for one pair in 2002 and one pair in 2003. Positive mean differences in juvenile salmon densities between ELJ and non‐ELJ units were observed in two of four years for all juvenile salmon, trout greater than 100 mm and juvenile Chinook salmon. Positive mean differences occurred in one of 4 years for juvenile coho salmon and trout less than 100 mm. The results suggest that ELJs are potentially useful for restoring juvenile salmon habitat in the Elwha River, Washington State, USA. Copyright © 2011 John Wiley & Sons, Ltd.  相似文献   

8.
The habitat use of subyearling Chinook salmon (Oncorhynchus tshawytscha) and coho salmon (Oncorhynchus kisutch) was examined in three tributaries of Lake Ontario. A total of 1781 habitat observations were made on Chinook salmon (698) and coho salmon (1083). During both spring and fall, subyearling coho salmon used pool habitat with abundant cover. During spring, principal component analysis revealed that water depth was the most important variable governing subyearling Chinook salmon habitat use. Substrate materials used by Chinook salmon in the spring and coho salmon in the fall were significantly smaller than were present on average within the study reaches. When the two species occurred sympatrically during spring they exhibited similar habitat selection. Although the habitat used by coho salmon in Lake Ontario tributaries was consistent with observations of habitat use in their native range, higher water velocities were less important to Chinook salmon than has previously been reported.  相似文献   

9.
Energy and resource fluxes between terrestrial and aquatic habitats of river flood plains can have reciprocal influences on food webs. Floodplain spring brooks may contain high densities (>1 m−2) of juvenile fish even when aquatic food resources appear to be low. A likely explanation is the allochthonous energy subsidy from riparian vegetation (e.g. leaves, invertebrates). To quantify this relationship in a rich salmon river ecosystem, we measured the temporal and spatial flux of terrestrial invertebrate and aquatic food resources in relation to diets of the two most abundant salmonid species in two spring brooks within early (<20 years) and late (>50 years) successional riparia on an expansive coastal flood plain of the Kol River, Kamchatka. Standing crops of benthic invertebrates and density of drifting invertebrates (dry mass) were fairly low (<4 g m−2 and <1 mg m−3, respectively) at both sites, but annual average terrestrial invertebrate input among sites was 22.2 ± 0.1 mg m−2 day−1. On average 68% of the coho salmon diet was of terrestrial origin, contrasting with 13% for Dolly Varden char at both sites, thus showing resource partitioning. Diet varied temporally and spatially with prey availability. Specifically, larvae of a weevil (Curculionidae) feeding on willow catkins were a dominant diet item for coho in the spring but only at the early successional site; Dolly Varden also ate these weevils but aquatic invertebrates continued to dominate their diets, further demonstrating behavioural segregation. The results show the importance of channel‐floodplain connectivity to management and conservation of salmon rivers. Copyright © 2009 John Wiley & Sons, Ltd.  相似文献   

10.
The river Lærdalselva, West Norway, was regulated in the autumn of 1974. Regulation led to an increase in winter flow and a decrease in summer flow in a section where there was natural production of salmon. A slight decrease in summer temperature was also recorded in the uppermost part of this section. No data existed on juvenile Atlantic salmon before regulation and the regulation effect on juvenile fish population parameters is therefore based on samples taken from adult salmon in the period 1969 to 1984. No differences in growth, smolt age, and smolt size which could be related to the regulation of the river were found. The smallest mean sizes of yearlings (0+) were found in 1964 and 1967. After regulation the mean size was never lower than before. Mean smolt age was between 3–1 and 3–9 years, but after regulation never exceeded that found before. River growth was slow, three year old smolts growing faster than four year old smolts. However, there were no differences in river growth before and after regulation.  相似文献   

11.
Low instream flows and high water temperatures are two factors limiting survival of native salmon in California's Shasta River. This study examines the potential to improve fish habitat conditions by better managing water quantity and quality using flow and water temperature simulation to evaluate potential restoration alternatives. This analysis provides a reasonable estimate of current and potential flows and temperatures for a representative dry year (2001) in the Shasta River, California. Results suggest restoring and protecting cool spring‐fed sources provides the most benefit for native salmon species from a broad range of restoration alternatives. Implementing a combination of restoration alternatives further improves instream habitat. Results also indicate that substituting higher quality water can sometimes benefit native species without increasing environmental water allocations. This study shows the importance of focusing on the limitations of specific river systems, rather than systematically increasing instream flow as a one size fits all restoration approach. Copyright © 2009 John Wiley & Sons, Ltd.  相似文献   

12.
Habitat use and habitat selection by young Atlantic salmon and brown trout were investigated by direct underwater observation. We sampled during winter and summer water temperatures (low: 3–7°C; high: 9–12°C) coinciding with low and high waterflows (12–20 and 60–80 m3 s?1), and during day and night in winter, and on six selected stations in the river. Observations of 396 salmon and 120 trout indicated a distinct seasonal pattern in behaviours and habitat selection. Feeding was the dominant behaviour at high water temperatures during summer. In winter, there was a diurnal pattern in behaviour; both species sheltered in interstitial spaces in the substrate during daylight, but during night held positions on or close to the substrate in slower flowing stream areas. Coarse substrate providing cover was therefore an important habitat factor during daylight at low water temperatures, while slow‐flowing water was important during night. Although spatial niche overlap was considerable both in summer and winter, salmon and trout segregated with respect to meso‐ and microhabitat selection, and relatively more at low temperatures. Both species changed their use of mesohabitats towards more slow‐flowing glide/flat habitats in winter. Irrespective of season, trout preferred in general more slow‐flowing water than salmon did, but the difference was more pronounced in winter. Salmon used a wider range of water depths and in particular water velocities, than did trout. Both species were less tolerant of high water velocities at low water temperatures. The seasonal and diurnal pattern in habitat selection reported have important implications for habitat research and habitat‐hydraulic modelling. Copyright © 2001 John Wiley & Sons, Ltd.  相似文献   

13.
The placement of boulder weirs is a popular method to improve fish habitat, though little is known about the effectiveness of these structures at increasing fish and biota abundance. We examined the effectiveness of boulder weir placement by comparing physical habitat, chemical and biotic metrics in 13 paired treatment (boulder weir placement) and control reaches in seven southwest Oregon watersheds in the summer of 2002 and 2003. Pool area, the number of boulders, total large woody debris (LWD) and LWD forming pools were all significantly higher in treatment than control reaches (p < 0.05). No differences in water chemistry (total N, total P, dissolved organic carbon) or macroinvertebrate metrics (richness, total abundance, benthic index of biotic integrity etc.) were detected. Abundance of juvenile coho salmon (Oncorhynchus kisutch) and trout (O. mykiss and O. clarki) were higher in treatment than control reaches (p < 0.05), while dace (Rhinichthys spp.; p < 0.09) were more abundant in control reaches and no significant difference was detected for young‐of‐year trout (p > 0.20). Both coho salmon and trout response to boulder weir placement were positively correlated with difference in pool area; p < 0.10), while dace and young‐of‐year trout response to boulder weir placement were negatively correlated with difference in LWD (p < 0.05). The placement of boulder weirs appears to be an effective technique for increasing local abundance of species that prefer pools (juvenile coho and trout >100 mm). Based on our results and previous studies on bedrock and incised channels, we suggest that the placement of boulder structures is a useful first step in attempting to restore these types of stream channels. Published in 2006 by John Wiley & Sons, Ltd.  相似文献   

14.
The effects of deep‐draft vessel traffic in confined riverine channels on shorelines and fish are of widespread concern. In the Pacific Northwest of the United States, wakes and subsequent beach run‐up from ships transiting the Lower Columbia River have been observed to strand juvenile salmon and other fish. As part of a before‐and‐after study to assess stranding effects that may be associated with channel deepening, we measured 19 variables from observations of 126 vessel passages at three low‐slope beaches and used multiple logistic regression to discern the significant factors influencing the frequency of stranding. Subyearling Chinook salmon were 82% of the fish stranded over all sites and seasons. Given a low‐slope beach, stranding frequencies for juvenile salmon were significantly related to river location, salmon density in the shallows, a proxy for ship kinetic energy, tidal height and two interactions. The beach types selected for our study do not include all the beach types along the Lower Columbia River so that the stranding probabilities described here cannot be extrapolated river‐wide. A more sophisticated modelling effort, informed by additional field data, is needed to assess salmon losses by stranding for the entire lower river. Such modelling needs to include river‐scale factors such as beach type, berms, proximity to navigation channel, and perhaps, proximity to tributaries that act as sources of outmigrating juvenile salmon. At both river and beach scales, no one factor produces stranding; rather interactions among several conditions produce a stranding event and give stranding its episodic nature. Published in 2010 by John Wiley & Sons, Ltd.  相似文献   

15.
The food habits of migrating juvenile steelhead (Salmo gairdneri), sockeye salmon (Oncorhynchus nerka), coho salmon (O. kisutch), and chinook salmon (O. tshawytscha) were identified from April through August 1984 at Bonneville Dam in the Columbia River. During the spring (April-June), the gammarid amphipods Corophium salmonis and C. spinicorne were the dominant prey for all species. Many insect taxa were also consumed, but in small quantities. Significant diet overlap occurred between all species during the spring due to the importance of Corophium. In summer (July-August), the importance of Corophium declined in the diet of subyearling chinook salmon and was replaced with Daphnia spp. and adult dipterans (primarily chironomids).  相似文献   

16.
Worldwide large rivers have been severely modified by human intervention. Many modifications result in disconnection of the river from floodplain and off‐channel habitats generally characterized by lower velocities and Copyright © 2016 John Wiley & Sons, Ltd. shallower depths relative to the main channel, conditions vital to many organisms. Extensive levees on the Middle Mississippi River (MMR) have cut off backwater systems and disconnected the river from 80% of its floodplain. However, the system is characterized by large side channels associated with islands. We examined a long term data set for differences in physical (e.g. depth and velocity) and water quality metrics (e.g. temperature, suspended solids, dissolved oxygen, chlorophyll, % organic matter) between the main and side channels of a 128‐km reach of the MMR. We compared variability between main and side channels using the coefficient of variation (COV). All metrics differed between habitats. Side channels were shallower with lower velocities and had greater mean and COV of % organic matter and more variable dissolved oxygen concentrations. Velocity, temperature, and suspended solids were similar in the spring. COVs were lowest in both habitats during the spring for all metrics except temperature and DO. Resource management in the MMR tends to focus on maintaining existing side channels because of the difficulty of working in the heavily used navigation channel. This study shows that these actions protect areas that function differently than the main channel for most of the year. However, our results also highlight the need for restoration activities aimed at restoring floodplain connectivity, especially during the spring. Copyright © 2016 John Wiley & Sons, Ltd.  相似文献   

17.
In winter, juvenile salmonids hide within the substrate during the day and emerge to feed on drifting invertebrates at night. In channelized streams, where the streambed heterogeneity has been artificially reduced, suitable microhabitats (low‐flow refugia) may be in short supply. Therefore, restoration of stream habitat by enhancement structures might improve the overwintering conditions of juvenile salmonids. We used a set of artificial streams to test whether individually‐marked juvenile brown trout of two age‐classes (age‐0 and age‐1 trout) loose mass during the winter differently in channelized and semi‐natural streams. Fish of both age‐classes lost mass early in the winter (November to January), but age‐0 fish in the channelized streams lost more of their initial mass than did the restored‐stream fish (ca. 10% vs. 2.5% on average, respectively). They then exhibited zero‐growth in both treatments in late winter (January to April), and by early spring (May), the channelized‐stream fish had completely caught up for their greater initial mass loss. In control tanks where the fish were fed continuously, age‐0 trout exhibited zero‐growth from November to January, then gaining mass constantly through the rest of the experiment. Significant time*treatment interaction was also detected for age‐1 trout, but all differences were caused by the faster growth of fish in the control tanks, whereas the two channel treatments did not differ significantly. The shortage of suitable sheltering sites in the channelized streams apparently intensified competition and caused greater initial mass loss in age‐0 trout. Furthermore, growth compensation exhibited by juvenile trout may have negative impacts on the long‐term fitness of individuals. Therefore, by increasing the amount of sheltering sites, in‐stream restoration may have potential to enhance the overwintering success of juvenile salmonids. Copyright © 2010 John Wiley & Sons, Ltd.  相似文献   

18.
Juvenile (post-smolt) coho salmon (Oncorhynchus kitsuch) were held in fresh water in the laboratory at 5, 10, 15, and 18°C for 8 weeks and fed freshly thawed, juvenile alewives (Alosa pseudoharengus) at rates equal to 1 and 2 % of their wet body weight/day, and also at the ad libitum or unrestricted ration rate. Most rapid growth in weight (1.2% wet body weight/day) occurred among fish fed the ad libitum ration at 15°C; growth was most rapid at about 10°C for fish fed the 2% ration (0.7% /day), and the 1% ration (0.1% /day). Gross conversion efficiency was highest at 10°C for all three ration levels. Gross body constituents and energy content of the test fish changed with temperature and ration during the study. Growth rate was positively related to lipid, energy content, and ration; lipid and energy content were positively related to water temperature; lipid, energy content, growth rate, ration, and water temperature were negatively related to water content; and protein was not related to any of the test variables. At the end of the study, water (68.7 to 76.4%) and lipid (3.5 to 10.4%) content were more variable than ash (1.8 to 3.1%), carbohydrate (0.1 to 1.9%), and protein (16.9 to 19.4%) content. Energy content of the fish increased with ration and was highest for each ration level at 15°C.  相似文献   

19.
Wintering cattle directly in the field creates potential concerns with water quality, as nutrients added from urine and fecal material over the winter can end up in runoff water, ground water and soil. In 2008/2009 an experiment was conducted to observe the effect of in-field winter feeding of cows on the nutrients in spring snowmelt run-off water. Low temperatures give little opportunity for organic N, urea and ammonium added in the urine and fecal matter to convert to nitrate, resulting in nitrate-N concentrations in snowmelt run-off water that were similar in the control and winter fed areas. Orthophosphate-P and ammonium-N concentrations were significantly elevated in run-off from the winter feed treatment basins compared to the controls. Surface soil sampled in the spring from the winter feeding site had higher soluble nitrate while soluble forms of phosphorus in the soil were lower compared to the fall soil samples. Caution should be used when utilizing in-field winter feeding systems so that the runoff water does not reach sensitive water bodies.  相似文献   

20.
In this paper, we use a system‐wide census of large river and floodplain habitat features to evaluate influences of valley form and land use on salmon habitats along 2,237 km of river in the Puget Sound region of Washington State, USA. We classified the study area by geomorphic process domains to examine differences in natural potential to form floodplain habitats among valley types, and by dominant land cover to examine land use influences on habitat abundance and complexity. We evaluated differences in aquatic habitat among strata in terms of metrics that quantify the length of main channels, side channels, braid channels, and area of wood jams. Among geomorphic process domains, habitat metrics standardized by main channel length were lowest in canyons where there is limited channel migration and less potential to create side channels or braids, and highest in post‐glacial and mountain valleys where island‐braided channels tend to form. Habitat complexity was lower in glacial valleys (generally meandering channels) than in post‐glacial valleys. Habitat abundance and complexity decreased with increasing degree of human influence, with all metrics being highest in areas classified as forested and lowest in areas classified as developed. Using multiple‐year aerial photography, we assessed the ability of our methods to measure habitat changes through time in the Cedar and Elwha Rivers, both of which have recent habitat restoration activity. We were able to parse out sources of habitat improvement or degradation through time, including natural processes, restoration, or development. Our investigation indicates that aerial photography can be an effective and practical method for regional monitoring of status and trends in numerous habitats.  相似文献   

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